#NEXUS [written Tue Apr 08 16:55:03 CEST 2025 by Mesquite version 3.81 (build 955) at DESKTOP-2PD9IQU/192.168.2.197] BEGIN TAXA; TITLE Taxa; DIMENSIONS NTAX=32; TAXLABELS Outgroup Abdobalistum Akromystax Coelodus_saturnus Flagellipinna Haqelpycnodus Iemanja Libanopycnodus Neoproscinetes Nursallia?_goedeli Nursallia_gutturosum Nursallia_veronae Ocloedus Oropycnodus Palaeobalistum Polazzodus_coronatus Polazzodus_gridellii Potiguara Proscinetes Pycnodus Scalacurvichthys Sigmapycnodus Stemmatodus Stenamara Sylvienodus Tepexichthys Tergestinia Thiollierepycnodus Turbomesodon_praeclarus Turbomesodon_relegans Turboscinetes Montenars_pycno ; END; BEGIN CHARACTERS; TITLE Character_Matrix; DIMENSIONS NCHAR=88; FORMAT DATATYPE = STANDARD RESPECTCASE GAP = - MISSING = ? SYMBOLS = " 0 1 2 3 4 5 6"; CHARSTATELABELS 1 'Body shape (as measured by ratio maximum body height/standard length)Body shape (as measured by ratio maxi- mum body height/standard length)' / 'discoid, 70-100%' 'intermediate, 40?70% ' 'fusiform, less than 40%' 'deep, more than 100% ', 2 Relative_position_of_dorsal_apex / between_the_skull_and_the_point_of_insertion_of_dorsal_fin_ apex_absent_ in_point_of_insertion_of_dorsal_fin_ in_skull_, 3 Dorsal_prominence / absent 'present, curved, dorsally oriented ', 4 Relative_position_of_ventral_apex / apex_absent before_point_of_insertion_of_anal_fin_ in_point_of_insertion_of_anal_fin_, 5 Mouth_gape / horizontal_or_subhorizontal_ inclined_, 6 Prognathism / 'absent (i.e., anterior part of mouth gape not distinctively projected, although the whole ethmoid region is hypertrophied)' 'present by elongation of mesethmoid, vomer and prearticular in the horizontal plane, marking a concavity in the anterior border of the head ' present_by_expansion_of_premaxilla_and_dentary_, 7 Caudal_pedicle / not_differentiated_ differentiated_, 8 Morphology_of_frontal_bones / narrow_above_the_orbit_ 'broad, expanded all throughout their length, nearly hemispheric in overall shape ', 9 Prefrontal_bones / absent_ present_, 10 Dermocranial_fenestra / absent_ present_, 11 Parietal_process / absent_ present, 12 'Extrascapular(s) fused to parietal' / no yes, 13 'Endocranium (supraoccipital bone)' / completely_covered_by_dermal_skull_in_lateral_view_ exposed_posteriorly_ largely_exposed_posteriorly, 14 'Postcephalic lacuna in endocranium (supraoccipital)' / 'supraoccipital not visible in late- ral view ' lacuna_absent lacuna_present, 15 Infraorbitals / 'reduced to tubular os- sifications around infraorbital sensory canal ' anterior_infraorbital_enlarged_ small_plates_ absent_, 16 Anterior_portion_of_infraorbital_sensory_canal / closely_surrounding_orbit_ descending_towards_ethmoid_region_, 17 Infraorbital_ornamentation / present_in_all_infraorbitals_ present_only_in_posteriormost_one absent_in_all_infraorbitals_, 18 Suborbitals / mosaic_of__small_plates absent_as_independent_ossifications_, 19 Preopercular_and_hyomandibular / 'preopercular single, hypertrophied; hyomandibular deep, unornamented' 'one high preopercular in close contact with small ornamented portion of hyomandibular, at same superficial level ' 'preopercular reduced, about as high as the ornamented portion of hyomandibular ' 'preopercular additionally reduced, clearly lower than ornamented portion of hyomandibular', 20 Condyle_in_articular_head_of_hyomandibular / absent_ present_, 21 Opercular_bone / reduced_ extremely_reduced, 22 Branchiostegal_rays / 'two, thin and separated' 'two, relatively large and in contact ' absent_ three_, 23 Morphology_of_premaxillary_and_dentary_teeth / 'robust, barely incisiform ' 'very flattened, fully incisiform' molariform_ incisiform_and_bifurcated, 24 Morphology_of_premaxilla / 'small, with long, slender process continuous with anterior border of main body of bone' 'large, rounded, horizontally expanded, with short, robust process emitted by posterior region of the bone', 25 Number_of_premaxillary_teeth / two three 'at least 8, arranged in at least two rows ' one, 26 'Morphology of maxilla (outline)' / ovoid_ reniform_ straight_oral_border elongated_oval oval_with_dorsal_notch_ 'axe- blade like ', 27 Crenulations_on_anteroventral_border_of_maxilla / absent_ present, 28 Morphology_of_vomerine_teeth / circular_to_subcircular_contour oval_contour reniform_contour_ triangular_contour_on_the_main_row 'oval, very elongated ', 29 Arrangement_of__vomerine_teeth_in_regular_rows / present_ 'absent anteriorly, present posteriorly', 30 Number_of__vomerine_tooth_rows / five three six, 31 Number_of_teeth_in_principal_vomerine_tooth_row / 10_or_more 8_or_9_ 7_or_less_, 32 Alternation_of_teeth_on_main_vomerine_tooth_row / absent present, 33 Number_of_dentary_teeth / four three two, 34 Morphology_of_prearticular_teeth / oval_contour_ circular_contour_ 'sigmoid to drop-shaped contour ' 'extremely elongated in contour, long axis perpendicular to row axis' 'oval, elongated, long axis of teeth coincident with row axis', 35 Arrangement_of_prearticular_teeth_in_regular_rows / present 'absent anteriorly, present posteriorly' 'absent anteriorly, present posteriorly' absent_, 36 Number_of__prearticular_tooth_rows / three two five_or_six, 37 Number_of_teeth_on_main_prearticular_tooth_row / 8_or_9_ 7_or_fewer 10_or_more, 38 Coronoid_process / 'high, straight dorsal border' 'high, club shaped', 39 Crenulations_in_vomerine_and_prearticular_teeth / 'occasionally present, weak' absent 'present in most teeth, strong', 40 Groove_on_vomerine_and_prearticular_teeth / absent present, 41 'Number of vertebrae (epichordal elements excluding those of the caudal endoskeleton)' / 30?34 35_or_more 25?29 24_or_fewer, 42 Neural_and_haemal_corresponding_arcocentra / not_surrounding_notochord surrounding_notochord_partially surrounding_notochord_completely, 43 Neural_and_haemal_adjacent_arcocentra / simple_contact complex_contact hyper_complex_contact expanded_and_imbricate, 44 Sagittal_flanges_on_neural_and_haemal_spines / 'anterior, small and short' 'anterior, large and long' anterior_and_posterior, 45 Number_of_autogenous_anterior_neural_spines / 10_or_more '7-9' 6_or_fewer, 46 Relative_length_of_last_neural_spine_not_supporting_precurrent_caudal_fin_rays / slightly_shorter_than_previous_ones less_than_half_as_long_as_preceding_spines vestigial, 47 Number_of_epichordal_elements_of_caudal_endoskeleton / 'primitive (?)' 6_to_8 4_or_5 3_or_fewer, 48 Relative_development_of__hypochordal_elements_of_caudal_endoskeleton / only_slightly_enlarged 'enlarged, plate-like' one_hypertrophied_element two_hypertrophied_elements, 49 Number_of_hypochordal_elements_of_caudal_endoskeleton / '9-11' '12-13' '6-8', 50 Diastema / absent present, 51 Cleithrum / 'curved, anteroventral limb subvertical, expanded' two_small_posterior_expansions_framing_a_shallow_notch_for_the_insertion_of_the_fin two_large_posterior_expansions_framing_a_deep_notch_for_the_insertion_of_the_fin_plus_anterior_elongation_of__the_anteroventral_limb_, 52 'Position of pelvic fins as ratio prepelvic distance/standard length' / '45-55%' more_than_55% less_than_45%, 53 'Position of dorsal fin as predorsal length/standard length' / '60%-69%' less_than_49% '50%-59%' '70%-79%', 54 Number_of_dorsal_axonosts / 30?39 40?49 50?59 60_or_more, 55 'Dorsal axonost not supporting lepidotrichium (free axonost)' / absent present, 56 Morphology_of_dorsal_and_anal_fins / 'primitive (?)' 'strip-like' falcate_to_acuminate sigmoid_outline rounded_in_center rounded_anteriorly 'extremely acuminate, higher anteriorly than it is long', 57 'Position of anal fin as preanal length/standard length ratio' / 70%?79% 50%?59% 60%?69% 80%?89%, 58 Number_of_anal_axonosts / 20?29 10?19 30?39 40?49 50_or_more 9_or_fewer, 59 Urodermals / one two absent, 60 Number_of_caudal_principal_fin_rays / 9_or_fewer '10-19' '20-25' '26-35' 36_ore_more, 61 Morphology_of_caudal_fin / distal_border_convex distal_border_concave double_emarginated vertical forked 'distal border vertical, straight', 62 Ossification_of_scales / complete_in_all_scales complete_in_ventral_scales_and_in_some_dorsal_scales 'complete in ventral scales, incomplete in dorsal scales' incomplete_in_all_scales, 63 Distribution_of_scales / only_abdominal_region whole_body_except_caudal_pedicle abdominal_region_plus_part_of__caudal_region whole_body, 64 Arrangement_of_scales / rows_in_same_direction rows_in_different_directions, 65 Ornamentation / tubercles ridges_ reticulation, 66 First_dorsal_ridge_scale / about_same_size_as_subsequent_ridge_scales larger_than_subsequent_ridge_scales, 67 'Morphology of second dorsal rid- ge scale' / similar_to_subsequent_dorsal_ridge_scales 'very long and large, with a central longitudinal ridge and a conspicuous anterior hook', 68 'Scutellum-like contour scales' / absent 'present, dorsal only' 'present, ventral only' 'present, dorsal and ventral', 69 Number_of_differentiated_dorsal_ridge_scales / 18_or_more 15_to_17 10_to_14 7_to_9, 70 Arrangement_of_dorsal_ridge_scales / dorsal_contour_scales_in_close_contact_with_each_other point_contact separated_from_each_other, 71 Number_of_spines_on_midline_of_dorsal_ridge_scales / 3_or_more midline_serrated 1_or_2 no_spines_on_dorsal_contour_scales, 72 Distribution_of_spines_on_midline_of_each_dorsal_ridge_scale / 'all along midline, or cente- red if only one spine present' no_spines_on_dorsal_contour_scales 'posterior region (at most two thirds) of midline' 'anterior region (at most two thirds) of midline', 73 Contact_of_spines_on_each_dorsal_ridge_scale / separated_from_each_other no_spines_on_dorsal_contour_scales in_contact_with_each_other, 74 Relative_size_of_anterior_and_posterior_spines_on_each_dorsal_ridge_scale / similar_in_size no_spines_on_dorsal_contour_scales spines_of_increasing_size_in_cephalocaudal_sense, 75 First_ventral_keel_scale / smaller_than_subsequent_scales larger_than_subsequent_scales, 76 Number_of_ventral_keel_scales / 22_or_more 18_to_21 15_to_17 10_to_14, 77 Arrangement_of_ventral_keel_scales / close_contact_with_each_other point_contact, 78 Number_of_spines_on_the_midline_of_ventral_keel_scales / 4_or_more 1_to_3 no_spines_on_ventral_keel_scales, 79 Distribution_of_spines_on_midline_of_ventral_keel_scales / 'all along midline, or centered if only one spine present' no_spines_on_ventral_keel_scales 'posterior region (at most two thirds) of midline', 80 Contact_of__spines_on_each_ventral_keel_scale / separated_from_each_other no_spines_on_ventral_keel_scales in_contact_with_each_other imbricate, 81 Relative_size_of_anterior_and_posterior_spines_on_each_ventral_keel_scale / spines_of_increasing_size_cephalocaudally no_spines_on_ventral_keel_scales, 82 Several_scales_attached_to_contour_scales / no yes, 83 'Number of post-cloacal ventral keel scales' / more_than_6 5_or_6 3_or_4 two, 84 Number_of_anterior_cloacal_modified_scales / mosaic_of_little_scales two one anterior_cloacal_scales_not_modified three, 85 'Number of posterior cloacal modi- fied scales' / mosaic_of_little_scales three two one 'no scales, posterior part of anal notch formed by a rib', 86 Bifid_scale_in_cloaca / absent 'present plus several comma-shaped scales', 87 'Post-cloacal notch' / absent present, 88 Supracloacal_scale / absent 'present, contacting only cloacal scales' 'present, contacting also non-differentiated scales adjacent to cloacal scales' ; MATRIX Outgroup 0000000000000000000000000000000000000000000000?00000000?000?0000000000000000000000000000 Abdobalistum 000?000??01?0011????0?00????0????000??01111121?2000?030114250321?10210200102021111?????? Akromystax 00010210111000???1200?212??401?00?0??000220110221000120512122200000020002213000300342000 Coelodus_saturnus 120?001?0?1?00???1????10???10000?3002?11010110?2010??302?3?32200??0?2231110??????0?????? Flagellipinna 02020010001?0020011?00300101????20???010??(2 3)22?????0?3?021????12020001000000?0????03??00? Haqelpycnodus 32021000011?0?11?11?01000??10???120???00000110111001200103?323000000220002???00000312000 Iemanja 1?0?0100001?00????110????30210?0?12??01021321021200?01??2323?2001??0?0????0??????0????0? Libanopycnodus 100010?0001?001??11?01100????????0?0?001200110?1??0101??0202?30001002010000??10000?????? Neoproscinetes 00011010001?000111110010010101111000111001111121200021?1222312001?00222302011020?0312?00 Nursallia?_goedeli ?????????????????????????????????????????221?1222??????2?41430110??????????????????????? Nursallia_gutturosum 0200001110?0001001100000000002020000010222122222000230224?23220110011????020??0003???0?? Nursallia_veronae 02000011?01?00??????0?000??001102?0?0?10222??2222000230224243120(0 1)10321311?031????02??10? Ocloedus 0202100001110001112000100??1000020000001200110211001200222122200000021022202002200322000 Oropycnodus 0201000001111201?12?1?100??10?2020001000221120130010230524232400(0 1 2)10131311103102200312210 Palaeobalistum 1100001????1002?0???0??0???00?10?0000?10???????100?00???1??4?0310?00112001010000?03????? Polazzodus_coronatus 100100100010113--130121035030020200000012101221200212202(0 2)(2 3)22440001102131110200000??23100 Polazzodus_gridellii 1001??10001021???12012???????????????0??2101221200212202222224000100213111031120003???00 Potiguara 02021000011?00?????????0???10000?000?00?2?01102100??13??231??????1????3111???00200?????? Proscinetes 0202100000110001111001100??10111200000100001102100012(1 2)1223131200110012311101102000311000 Pycnodus 1000001001101211013000100100001020002010221122130010120113232400(0 2)10131311103102000323100 Scalacurvichthys 100110000010?????10?0?100??10??0200??00030011120000?0?0?0??30400(0 1)??0??3111??012000???000 Sigmapycnodus 1???0??0??1?00??????????????????0????1??1111?0????0?1???1????30????????????????????????? Stemmatodus 10001000001?0001112000100200001020002000200120200000001122121300200020022203000200324000 Stenamara 30111000001?0001???00??0?????????00100?0200110302002001?2012?200?000200022010???00223000 Sylvienodus 110110100011113--120121035110?102????0203201102100110006(0 2)2022400(0 2)00021102002002200322100 Tepexichthys 020110000111000121100?1014000200100001100001112100002(0 2)021(0 2)231200100010311101000200113000 Tergestinia 130110100010113--13?12103??1001022001021210120120010220122215400200121311103010??0323100 Thiollierepycnodus 02020010001?0001111001300000??0000????00100110?0000201122312232010?00(0 2)3111?0012??00??00? Turbomesodon_praeclarus 000110000011000111100?100101010020001101000110302000000420120200200011020201000200123002 Turbomesodon_relegans 020010000011000111100?1001010?00200010010001103000000(0 1)132212220020001(1 2)020201000200123001 Turboscinetes 000000000011000111100310110100101000?011(0 2)0011011000031122(0 2)0123001100(0 1)231110111111021(2 3)000 Montenars_pycno 12000010011?1201013000100501011023000011210120230010220123?12400000021311102000000333100 ; END; BEGIN ASSUMPTIONS; TYPESET * UNTITLED = unord: 1- 88; END; BEGIN MESQUITECHARMODELS; ProbModelSet * UNTITLED = 'Mk1 (est.)': 1- 88; END; BEGIN NOTES; TEXT TAXON = 4 CHARACTER = 69 TEXT = 2; END; Begin MESQUITE; MESQUITESCRIPTVERSION 2; TITLE AUTO; tell ProjectCoordinator; timeSaved 1744124103942; getEmployee #mesquite.minimal.ManageTaxa.ManageTaxa; tell It; setID 0 1987887432145750149; tell It; setDefaultOrder 0 1 2 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 33 31; attachments ; endTell; endTell; getEmployee #mesquite.charMatrices.ManageCharacters.ManageCharacters; tell It; setID 0 4513173557149214512; mqVersion 381; checksumv 0 3 671352389 null getNumChars 88 numChars 88 getNumTaxa 32 numTaxa 32 short true bits 127 states 127 sumSquaresStatesOnly 34014.0 sumSquares 34014.0 longCompressibleToShort false usingShortMatrix true NumFiles 1 NumMatrices 1; mqVersion; endTell; getWindow; tell It; suppress; setResourcesState false false 100; setPopoutState 400; setExplanationSize 0; setAnnotationSize 0; setFontIncAnnot 0; setFontIncExp 0; setSize 1064 945; setLocation 5 12; setFont SanSerif; setFontSize 10; getToolPalette; tell It; endTell; desuppress; endTell; getEmployee #mesquite.minimal.ManageTaxa.ManageTaxa; tell It; showTaxa #1987887432145750149 #mesquite.lists.TaxonList.TaxonList; tell It; setTaxa #1987887432145750149; getWindow; tell It; useTargetValue off; setTargetValue ; newAssistant #mesquite.lists.TaxonListCurrPartition.TaxonListCurrPartition; newAssistant #mesquite.lists.DefaultTaxaOrder.DefaultTaxaOrder; setExplanationSize 30; setAnnotationSize 20; setFontIncAnnot 0; setFontIncExp 0; setSize 964 873; setLocation 5 12; setFont SanSerif; setFontSize 10; getToolPalette; tell It; setTool mesquite.lists.TaxonList.TaxonListWindow.ibeam; endTell; endTell; showWindow; getEmployee #mesquite.lists.ColorTaxon.ColorTaxon; tell It; setColor Red; removeColor off; endTell; getEmployee #mesquite.lists.TaxonListAnnotPanel.TaxonListAnnotPanel; tell It; togglePanel off; endTell; endTell; endTell; getEmployee #mesquite.charMatrices.BasicDataWindowCoord.BasicDataWindowCoord; tell It; showDataWindow #4513173557149214512 #mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindowMaker; tell It; getWindow; tell It; getTable; tell It; rowNamesWidth 109; endTell; setExplanationSize 30; setAnnotationSize 20; setFontIncAnnot 0; setFontIncExp 0; setSize 964 873; setLocation 5 12; setFont SanSerif; setFontSize 10; getToolPalette; tell It; setTool mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindow.ibeam; endTell; setTool mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindow.ibeam; colorCells #mesquite.charMatrices.NoColor.NoColor; colorRowNames #mesquite.charMatrices.TaxonGroupColor.TaxonGroupColor; colorColumnNames #mesquite.charMatrices.CharGroupColor.CharGroupColor; colorText #mesquite.charMatrices.NoColor.NoColor; setBackground White; toggleShowNames on; toggleShowTaxonNames on; toggleTight off; toggleThinRows off; toggleShowChanges on; toggleSeparateLines off; toggleShowStates on; toggleReduceCellBorders off; toggleAutoWCharNames on; toggleAutoTaxonNames off; toggleShowDefaultCharNames off; toggleConstrainCW on; toggleBirdsEye off; toggleColorOnlyTaxonNames off; toggleShowPaleGrid off; toggleShowPaleCellColors off; toggleShowPaleExcluded off; togglePaleInapplicable on; togglePaleMissing off; toggleShowBoldCellText off; toggleAllowAutosize on; toggleColorsPanel off; toggleDiagonal on; setDiagonalHeight 80; toggleLinkedScrolling on; toggleScrollLinkedTables off; endTell; showWindow; getWindow; tell It; forceAutosize; endTell; getEmployee #mesquite.charMatrices.AlterData.AlterData; tell It; toggleBySubmenus off; endTell; getEmployee #mesquite.charMatrices.ColorByState.ColorByState; tell It; setStateLimit 9; toggleUniformMaximum on; endTell; getEmployee #mesquite.charMatrices.ColorCells.ColorCells; tell It; setColor Red; removeColor off; endTell; getEmployee #mesquite.categ.StateNamesEditor.StateNamesEditor; tell It; makeWindow; tell It; getTable; tell It; rowNamesWidth 238; endTell; setExplanationSize 30; setAnnotationSize 20; setFontIncAnnot 0; setFontIncExp 0; setSize 964 873; setLocation 5 12; setFont SanSerif; setFontSize 10; getToolPalette; tell It; setTool mesquite.categ.StateNamesEditor.StateNamesWindow.ibeam; endTell; setActive; rowsAreCharacters on; toggleConstrainChar on; toggleConstrainCharNum 3; togglePanel off; toggleSummaryPanel off; endTell; showWindow; endTell; getEmployee #mesquite.categ.StateNamesStrip.StateNamesStrip; tell It; showStrip off; endTell; getEmployee #mesquite.charMatrices.AnnotPanel.AnnotPanel; tell It; togglePanel off; endTell; getEmployee #mesquite.charMatrices.CharReferenceStrip.CharReferenceStrip; tell It; showStrip off; endTell; getEmployee #mesquite.charMatrices.QuickKeySelector.QuickKeySelector; tell It; autotabOff; endTell; getEmployee #mesquite.charMatrices.SelSummaryStrip.SelSummaryStrip; tell It; showStrip off; endTell; getEmployee #mesquite.categ.SmallStateNamesEditor.SmallStateNamesEditor; tell It; panelOpen true; endTell; endTell; endTell; getEmployee #mesquite.charMatrices.ManageCharacters.ManageCharacters; tell It; showCharacters #4513173557149214512 #mesquite.lists.CharacterList.CharacterList; tell It; setData 0; getWindow; tell It; useTargetValue off; setTargetValue ; newAssistant #mesquite.lists.DefaultCharOrder.DefaultCharOrder; newAssistant #mesquite.lists.CharListInclusion.CharListInclusion; newAssistant #mesquite.lists.CharListPartition.CharListPartition; newAssistant #mesquite.parsimony.CharListParsModels.CharListParsModels; setExplanationSize 30; setAnnotationSize 20; setFontIncAnnot 0; setFontIncExp 0; setSize 964 873; setLocation 5 12; setFont SanSerif; setFontSize 10; getToolPalette; tell It; setTool mesquite.lists.CharacterList.CharacterListWindow.ibeam; endTell; endTell; showWindow; getEmployee #mesquite.lists.CharListAnnotPanel.CharListAnnotPanel; tell It; togglePanel off; endTell; endTell; endTell; endTell; end;