RIVISTA ITALIANA DI PALEONTOLOGIA E STRATIGRAFIA https://riviste.unimi.it/index.php/RIPS <p>The Rivista Italiana di Paleontologia e Stratigrafia is a <strong>free of charge</strong> <strong>OPEN ACCESS </strong>peer reviewed journal.</p> <p>RIPS publishes original contributions on all aspects of palaeontology and stratigraphy; all papers are written in English and are reviewed by international experts.</p> <p>The Journal is currently indexed and abstracted by AGI, ISI, Current Contents, Georef, Geological Abstract and SciSearch.</p> <p><strong>RIPS is included in the Unimi Journals project (<a href="https://riviste.unimi.it/">riviste.unimi.it</a>) and it is hosted on <a href="http://pkp.sfu.ca/?q=ojs">OJS</a> platform.</strong></p> <p><strong>OPEN ACCESS POLICY</strong>: RIPS is published under a <a href="http://creativecommons.org/licenses/by-sa/4.0/" rel="license">Creative Commons Attribution - Share alike 4.0 International License</a>.</p> <p>Preprint and postprint version of the articles can be archived anywhere with the same license.</p> <p> </p> <p><em><strong>Web of Science Journal Citation Reports 2023</strong>: Journal Impact Factor 2021= <strong>1.6</strong>, 5-Year Impact Factor= <strong>1.7</strong></em></p> <p><em><strong>Scopus CiteScore 2023</strong>= <strong>3.6</strong></em></p> <p class="evidenza"> </p> <p class="evidenza"><strong>Editors-in-chief</strong></p> <p>Lucia Angiolini, Università degli Studi di Milano, Italy<br />Fabrizio Berra, Università degli Studi di Milano, Italy</p> <p><strong>Editorial Office</strong></p> <p>Cristina Lombardo, Università degli Studi di Milano, Italy</p> <p class="evidenza"><strong>Section Editors</strong></p> <p>David M. Alba, Universitat Autònoma de Barcelona, Spain<br />Francesca Bosellini, Università degli Studi di Modena e Reggio Emilia, Italy<br />Giorgio Carnevale, Università degli Studi di Torino, Italy<br />Silvia Gardin, CNRS, France<br />Luca Giusberti, Università degli Studi di Padova, Italy<br />Annette Götz, Georg-August-Universität Göttingen, Germany<br />Joachim Haug, Ludwig-Maximilians-Universität München, Germany<br />Ilaria Mazzini, Consiglio Nazionale delle Ricerche, Roma, Italy<br />Giovanni Muttoni, Università degli Studi di Milano, Italy<br />Maria Rose Petrizzo, Università degli Studi di Milano, Italy<br />Isabella Raffi, Università degli Studi "G. D'Annunzio" Chieti Pescara, Italy<br />Silvio Renesto, Università dell'Insubria, Italy<br />Lorenzo Rook, Università degli Studi di Firenze, Italy<br />Daniele Scarponi, Università di Bologna, Italy<br />Lars Werdelin, Swedish Museum of Natural History, Sweden</p> <p> </p> <p><strong>Print subscriptions to Rivista Italiana di Paleontologia e Stratigrafia are handled by the publisher <a href="http://www.ledizioni.it/">Ledizioni</a> The Innovative LediPublishing company and subscriptions are managed by <a href="http://www.internationalbookseller.com">Libreria Ledi International Bookseller</a></strong></p> <p>Email: <a href="mailto:riviste@internationalbookseller.com">riviste@internationalbookseller.com</a></p> <p> </p> <p><strong>RIPS</strong> on Facebook: <a href="https://www.facebook.com/RIPS15/" target="_blank" rel="noopener">https://www.facebook.com/RIPS15/</a></p> <p> </p> Milano University Press en-US RIVISTA ITALIANA DI PALEONTOLOGIA E STRATIGRAFIA 0035-6883 <p>The journal allow the author(s) to hold the copyright without restrictions.</p> FIRST REPORT ON THE UPPERMOST PERMIAN OSTRACODS FROM THE MASORE SECTION (EXTERNAL DINARIDES), SLOVENIA https://riviste.unimi.it/index.php/RIPS/article/view/21186 <p>The ostracod assemblages from the Upper Permian and Permian-Triassic transitional strata of the Masore section in the External Dinarides (Slovenia) were studied. Altogether 13 genera and 20 species of the orders Palaeocopida, Platycopida and Podocopida are identified and illustrated. All recovered ostracods belong to shallow marine taxa. The older fauna from the uppermost Permian and the younger fauna from the Permian-Triassic boundary strata are distinguished. The obtained fauna from the Permian Bellerophon Formation corresponds to the “benthonic ostracod Eifelian Mega-assemblage” occupying the shelf sea floors. This study presents the first report of ostracod faunas from the uppermost Permian and Permian-Triassic boundary interval of Slovenia. The recovered ostracod fauna display a distinct faunal change and yield important paleobiogeographic implications as it reveals similarity with stratigraphical equivalent faunas from some other neighboring localities in the western Paleotethys, such as Bulla in Italy, Komirić in Serbia, and the Bükk Mountains in Hungary.</p> TEA KOLAR-JURKOVŠEK EWA OLEMPSKA BOGDAN JURKOVŠEK Copyright (c) 2024 TEA KOLAR-JURKOVŠEK, EWA OLEMPSKA , BOGDAN JURKOVŠEK https://creativecommons.org/licenses/by-sa/4.0 2024-05-16 2024-05-16 130 2 10.54103/2039-4942/21186 REVISITING THE CRETACEOUS LUNGFISH <em>ATLANTOCERATODUS IHERINGI</em> (AMEGHINO 1898) FROM THE MATA AMARILLA FORMATION (ARGENTINA) WITH COMMENTS ON TOOTH PLATES HISTOLOGY https://riviste.unimi.it/index.php/RIPS/article/view/21646 <p><em>Atlantoceratodus iheringi</em> (Ameghino 1898) from Argentine territory is restudied based on its known tooth plates and newly discovered material, which includes previously unknown skull roof bones and vomerine tooth plates. The latter represent the first records of such elements from the Mesozoic era in South America. The comparative morphological analysis reveals its distinctiveness from other dipnoans, and offers valuable data for future systematic and phylogenetic research. The pterygopalatine tooth plates display narrow-based first denticulations and lack anterior wear facets, with the inner angle positioned at the level of the second denticulation. Similarly, the prearticular tooth plates feature straight mediolingual edges, and a wide-based first denticulation without sinuosities at the tip. Histological sections are performed and analyzed in detail for the first time. <em>A. iheringi</em> presents in this aspect distinctive features such as: large-lumen denteons tending to cluster together, circumdenteonal dentine arranged in a double band (an inner birefringent and an outer monorefringent), and a disordered interdenteonal dentine, with birefringence surrounding denteons and areas with monorefringence. <em>A. iheringi</em> exhibits histological structure closer to Mesozoic and Cenozoic dipnoans than Paleozoic, especially resembling the disposition observed in the Upper Cretaceous Patagonian species <em>Metaceratodus baibianorum</em>. The wide distribution of features designated as diagnostic for <em>Atlantoceratodus</em> is discussed.</p> KAREN M. PANZERI Copyright (c) 2024 KAREN M. PANZERI https://creativecommons.org/licenses/by-sa/4.0 2024-05-16 2024-05-16 130 2 10.54103/2039-4942/21646 NEW ACROPOMATIFORM FOSSILS FROM THE UPPER KUEICHULIN FORMATION (LOWER PLIOCENE), NORTHERN TAIWAN https://riviste.unimi.it/index.php/RIPS/article/view/22639 <p>The Indo-West Pacific is home to a diverse array of modern fish species, and the study of their fossils can provide valuable insights into the origins and evolution of this biodiversity. This report presents rare remains of five individuals of acropomatiform fish, including a semi-articulated skeleton, recovered from the Upper Kueichulin Formation (Lower Pliocene) in northern Taiwan. The fossils are preserved in the form of nodules discovered at the locality known for abundant marine crustaceans and echinoderms. These specimens can be assigned to the family Stereolepididae, based on morphology of the opercle with the presence of two spines, a preopercle characterized by serrations on the posterior margin and denticulations on the ventral margin, and a distinctive skull architecture. Differences in skull morphology distinguish these specimens from other species within the genus and thus are described as Stereolepis arcanum n. sp. Significantly, the species represents the oldest known record of the family globally and stands as the second known fossil representative. This discovery not only enhances our understanding of the palaeobiodiversity of fish fauna in the Indo-West Pacific during the Neogene, but also highlights the importance of further research efforts in this area.</p> TOMÁŠ PŘIKRYL CHIEN-HSIANG LIN CHIA-HSIN HSU SHIH-WEI LEE Copyright (c) 2024 TOMÁŠ PŘIKRYL, CHIEN-HSIANG LIN, CHIA-HSIN HSU, SHIH-WEI LEE https://creativecommons.org/licenses/by-sa/4.0 2024-05-22 2024-05-22 130 2 10.54103/2039-4942/22639 PENNSYLVANIAN CONODONTS AND MICROFACIES FROM NORTHEASTERN MEXICO (TAMATÁN GROUP, CIUDAD VICTORIA BLOCK) https://riviste.unimi.it/index.php/RIPS/article/view/22324 <p>The biostratigraphic analysis of the Paleozoic sedimentary successions in Mexico is crucial for understanding their age, depositional setting, and paleogeographic implications. This study examined conodonts and microfacies of calcareous strata within the Del Monte Formation of the Tamatán Group in northeastern Mexico.</p> <p>Field observations have led to the identification of three units within the Del Monte Formation: Unit 1, characterized by calcareous sandstone, bioclastic limestone, shale, occasional conglomerate, and dolomitized beds; Unit 2, consisting of conglomerate and/or breccia associated with debris flows; and Unit 3 including turbiditic sandstone and shale.</p> <p>Conodonts were only recognized in Unit 1, with such genera as <em>Idiognathoides</em>, <em>Neognathodus</em>, <em>Idiognathodus</em> and less frequently, <em>Streptognathodus</em>, <em>Adetognathus</em>, and <em>Diplognathodus</em>. The faunal associations include <em>Idiognathoides</em> cf. <em>Id. corrugatus</em> Harris &amp; Hollingsworth, 1933<em>, Idiognathoides sulcatus sulcatus</em> Higgins &amp; Bouckaert, 1968<em>, Idiognathoides convexus </em>(Ellison &amp; Graves, 1941)<em>, Idiognathoides asiaticus </em>Nigmadganov &amp; Nemyrovskaya, 1992<em>, Neognathodus</em> cf. <em>N. atokaensis</em> Grayson, 1984<em>, Neognathodus</em> aff. <em>bothrops </em>Merrill, 1972<em>, Neognathodus nataliae </em>Alekseev &amp; Gerelzezeg, 2001 (in Alekseev &amp; Goreva 2001)<em>, Streptognathodus parvus </em>Dunn, 1966<em>, Idiognathodus incurvus </em>Dunn, 1966, and <em>Idiognathodus </em>aff. <em>delicatus </em>Gunnell, 1931, suggesting Morrowan and Atokan (Pennsylvanian) ages for the dolomitic limestone and grainstone beds.</p> <p>Five types of microfacies (MF) and several marine facies zone (FZ) have been detected: MF1-bioclastic grainstone and MF2-wackestone deposited at the platform margin (FZ 6 and FZ 7), MF3-dolomitized wackestone representing an inner restricted platform (FZ 8), MF4-mudstone from a deeper water zone (FZ 4), and MF5-siliciclastic beds with bioclasts indicating a high-energy barrier associated with a platform margin (FZ 6). The findings confirm a shallow to deep platform and imply a connection among Mexican peri-Gondwanan basins, including the Tamatán Basin, indicating similarities in sedimentary facies and/or depositional environments linked to the Rheic Ocean during the Early to Middle Pennsylvanian.</p> JUAN MOISÉS CASAS-PEÑA PILAR NAVAS-PAREJO UWE JENCHEN JUAN ALONSO RAMÍREZ-FERNÁNDEZ Copyright (c) 2024 JUAN MOISÉS CASAS-PEÑA, PILAR NAVAS-PAREJO, UWE JENCHEN, JUAN ALONSO RAMÍREZ-FERNÁNDEZ https://creativecommons.org/licenses/by-sa/4.0 2024-06-06 2024-06-06 130 2 10.54103/2039-4942/22324 NEW INSIGHTS ON THE ENIGMATIC OTTERS FROM THE LATE MIOCENE OF TUSCANY: <em>TYRRHENOLUTRA MAREMMANA</em> NOV. COMB. (LUTRINAE, MUSTELIDAE, CARNIVORA), WITH A PHYLOGENY OF BUNODONT OTTERS https://riviste.unimi.it/index.php/RIPS/article/view/22479 <p>Otters, subfamily Lutrinae (Mustelidae), are predators that thrive in aquatic environments, and have a semiaquatic to fully aquatic lifestyle. While modern otters are found across the globe, their fossil history is limited, primarily documented by fragmented dentognathic remains. In this work, we report on and describe new dental remains of a medium-sized Lutrinae from the Late Miocene of Baccinello and Casteani (Tuscany), improving the poor fossil record of the subfamily in Europe. <em>Tyrrhenolutra helbingi </em>Hürzeler, 1987 is a poorly known species, described on some dental remains from the Baccinello mines and a few other localities. Considering its peculiar morphology, it has often been overlooked in the scientific literature. Description of the material stored at the Naturhistorisches Museum, Basel and Museo di Geologia e Paleontologia dell’Università di Firenze offers new insights on the intraspecific variability and dental morphology of this taxon. The comparative study of these findings highlights several morphological and biometrical similarities between <em>T. helbingi </em>and <em>Paludolutra maremmana </em>Hürzeler, 1987, which allow us to redefine these taxa as <em>Tyrrhenolutra maremmana </em>nov. comb. We carried out a total-evidence cladistic analysis seeking support for morphometric and morphological similarities between <em>T. </em>“<em>helbingi</em>” and “<em>P.</em>” <em>maremmana</em>. The resulting parsimony analyses support the synonymy, excluding a close relationship of <em>T.</em> <em>maremmana</em> with <em>Paludolutra </em>of Italy and Spain, as previously thought. Furthermore, we confirm a long-forgotten hypothesis identifying <em>Limnonyx </em>Crusafont Pairó, 1950 as the possible continental ancestor of <em>Tyrrhenolutra</em>.</p> ANDREA FAGGI SAVERIO BARTOLINI-LUCENTI LORENZO ROOK Copyright (c) 2024 ANDREA FAGGI, SAVERIO BARTOLINI-LUCENTI, LORENZO ROOK https://creativecommons.org/licenses/by-sa/4.0 2024-06-13 2024-06-13 130 2 10.54103/2039-4942/22479 THE FIRST RECORD OF THE ORECTOLOBIFORM SHARK GENUS <em>CEDERSTROEMIA</em> (ELASMOBRANCHII, ORECTOLOBIDAE) IN ASIA (KASHIMA FORMATION, UPPER CRETACEOUS; OYUBARI AREA, HOKKAIDO, JAPAN) https://riviste.unimi.it/index.php/RIPS/article/view/22195 <p>We described fossil teeth assigned to the orectolobiform shark genus, <em>Cederstroemia</em> collected from the tuffaceous sandstone of the Kashima Formation (Santonian) in the Oyubari area, Hokkaido, Japan. This is the first report of fossil remains of <em>Cederstroemia</em> from Asia. Due to the small number of specimens and their poor preservation, we left the studied material in open nomenclature as <em>Cederstroemia</em> sp. At the same time, the studied material may belong to a new previously undescribed morphology of <em>Cederstroemia</em>. The occurrence of <em>Cederstroemia</em> fossils in Japan indicates that this genus dispersed to the northwestern part of the Pacific Ocean in the Late Cretaceous, which considerably extends the range and previously known habitat of this genus.</p> MASAHIKO KANEKO SERGEY V. SOLONIN Copyright (c) 2024 MASAHIKO KANEKO, SERGEY V. SOLONIN https://creativecommons.org/licenses/by-sa/4.0 2024-06-14 2024-06-14 130 2 10.54103/2039-4942/22195 AN UNEXPECTED SHORT TOOTH REPLACEMENT CYCLE PERIOD IN <em>MARESAURUS COCCAI</em> (PLESIOSAURIA; RHOMALEOSAURIDAE) FROM THE BAJOCIAN OF ARGENTINEAN PATAGONIA https://riviste.unimi.it/index.php/RIPS/article/view/22114 <p>The Rhomaleosauridae were a clade of Jurassic plesiosaurians, characterized by triangular skulls, elongated preorbital region, marked premaxillary-maxillary constriction, and intermediate body proportions between pliosauromorphs and plesiosauromorphs. Despite recent progress in the study of dental replacement in plesiosaurians, the replacement features in rhomaleosaurids have not been studied. Here, the dental features of the rhomaleosaurid <em>Maresaurus cocca</em><em>i</em> are described and analyzed based on the holotype specimen. Regarding symmetry, it was determined that <em>M. coccai </em>shows symmetrical replacement in the maxillary-premaxillary and asymmetrical replacement in the dentary. Additionally, the tooth replacement cycle period (TRCP) of two alveoli was determined for the dental series, with the exception of the anterior part of the left mandibular ramus, which presents an TRCP of three, an asymmetry considered here as teratology. This result indicates that the replacement cycle period (TRCP) of the two alveoli would correspond to a primitive character for Plesiosauria, observing an increase of TRCP from two to three alveoli in taxa comparatively more derived within Pliosauridae.</p> GONZALO MATELO MIRCO JOSÉ PATRICIO O’GORMAN ZULMA GASPARINI Copyright (c) 2024 GONZALO MATELO MIRCO, JOSÉ PATRICIO O’GORMAN, ZULMA GASPARINI https://creativecommons.org/licenses/by-sa/4.0 2024-06-17 2024-06-17 130 2 10.54103/2039-4942/22114 DIGITAL INVESTIGATION OF LAMNIFORM SHARK VERTEBRAE FROM THE SIBILLINI MTS. (NORTHERN APENNINES, ITALY) https://riviste.unimi.it/index.php/RIPS/article/view/22571 <p>During the sampling of a stratigraphic section along the shore of the Fiastra Lake (Carg Project - Sheet 313 “Camerino” of the Geological Map of Italy at 1:50 000 scale), a small rock boulder with partially exposed bony material was discovered at the base of a small cliff at the northern termination of the Sibillini Mts. In this area, the classical facies of Umbria-Marche stratigraphic succession are well exposed. The Oligocene-Miocene portion of the succession is represented by the ~200 m-thick Scaglia Cinerea Formation, passing upwards to the ~100 m-thick Bisciaro Formation. The microfossil assemblage has allowed the specimen to be constrained to the lower Burdigalian. The skeletal remains were examined using a CT-SCAN, a non-invasive method that has proven to be highly performing. The analysis revealed some articulated vertebrae, deformed by lithostatic compaction, which are attributed to a shark of the Order Lamniformes. Subsequently, the vertebrae were digitally isolated, extracted from the surrounding matrix, and rendered into three-dimensional prints. Through digital retro-deformation, the body length of the lamniform shark was estimated to be approximately 4 metres. Further considerations on the vertebrae allowed us to infer that the studied shark had similarities to either <em>Isurus oxyrinchus </em>Rafinesque, 1810 or <em>Carcharodon carcharias</em> Linnaeus, 1758<em>. </em>The development of a dead-fall microbial community likely facilitated the preservation of the vertebrae. The studied specimen represents the first occurrence of a lamniform shark in the Lower Miocene of the Umbria-Marche Domain and represents one of the very rare recorded occurrences of lamniforms from the Lower Miocene of Italy.</p> ALFIO MOSCARELLA MARCO ROMANO LORENZO CONSORTI ANGELO CIPRIANI GABRIELE BINDELLINI GIUSEPPE MARRAMÀ GIORGIO CARNEVALE ADELE GARZARELLA MARIA LETIZIA PAMPALONI PAOLO CITTON FERDINANDO SPANO' CHIARA D'AMBROGI CRISTINA MURARO ERNESTO PAOLO PRINZI GIUDITTA RADEFF GINO ROMAGNOLI SIMONE FABBI Copyright (c) 2024 ALFIO MOSCARELLA, MARCO ROMANO, LORENZO CONSORTI, ANGELO CIPRIANI, GABRIELE BINDELLINI, GIUSEPPE MARRAMÀ, GIORGIO CARNEVALE, ADELE GARZARELLA, MARIA LETIZIA PAMPALONI, PAOLO CITTON, FERDINANDO SPANO', CHIARA D'AMBROGI, CRISTINA MURARO, ERNESTO PAOLO PRINZI, GIUDITTA RADEFF, GINO ROMAGNOLI, SIMONE FABBI https://creativecommons.org/licenses/by-sa/4.0 2024-06-24 2024-06-24 130 2 10.54103/2039-4942/22571 THE CARNIVORAN RECORD FROM THE NEOGENE OF EASTERN ROMANIA https://riviste.unimi.it/index.php/RIPS/article/view/22194 <p>The present work deals with the carnivorans found in five Neogene localities in eastern Romania: the Vallesian locality of Păun, the Turolian localities of Pogana 1 and Creţeşti 1, and the Ruscinian localities of Bereşti and Măluşteni. The presented material includes cranial and postcranial specimens, some of which are herein described for the first time. This study includes a re-description and re-depiction of the type material of <em>Lutra rumana</em> Simionescu, 1922 and <em>Promephitis</em> <em>malustenensis</em> Simionescu, 1930, in order to clarify the taxonomic position of these two enigmatic taxa. The carnivoran faunal lists of all five localities are reviewed offering interesting insights into the temporospatial range of the discovered forms. Most localities have only yielded a limited number of species, except from Măluşteni, which shows evidence of at least nine different coexisting forms. Notable occurrences in terms of biostratigraphy include <em>Paludolutra </em>sp. in Pogana 1, <em>Protictitherium crassum</em> (Depéret, 1892) in Creţeşti 1, and a phocid from Măluşteni. This review aims to aid the understanding of the Neogene ecosystems in the north of the Paratethys in terms of faunal contents, biostratigraphy and palaeoecology.</p> <p> </p> <p> </p> NIKOLAOS KARGOPOULOS ALBERTO VALENCIANO PANAGIOTIS KAMPOURIDIS ŞTEFAN VASILE LAURENŢIU URSACHI BOGDAN RĂŢOI Copyright (c) 2024 NIKOLAOS KARGOPOULOS, ALBERTO VALENCIANO, PANAGIOTIS KAMPOURIDIS, ŞTEFAN VASILE, LAURENŢIU URSACHI, BOGDAN RĂŢOI https://creativecommons.org/licenses/by-sa/4.0 2024-06-28 2024-06-28 130 2 10.54103/2039-4942/22194