THE CARNIVORAN RECORD FROM THE NEOGENE OF EASTERN ROMANIA
DOI:
https://doi.org/10.54103/2039-4942/22194Abstract
The present work deals with the carnivorans found in five Neogene localities in eastern Romania: the Vallesian locality of Păun, the Turolian localities of Pogana 1 and Creţeşti 1, and the Ruscinian localities of Bereşti and Măluşteni. The presented material includes cranial and postcranial specimens, some of which are herein described for the first time. This study includes a re-description and re-depiction of the type material of Lutra rumana Simionescu, 1922 and Promephitis malustenensis Simionescu, 1930, in order to clarify the taxonomic position of these two enigmatic taxa. The carnivoran faunal lists of all five localities are reviewed offering interesting insights into the temporospatial range of the discovered forms. Most localities have only yielded a limited number of species, except from Măluşteni, which shows evidence of at least nine different coexisting forms. Notable occurrences in terms of biostratigraphy include Paludolutra sp. in Pogana 1, Protictitherium crassum (Depéret, 1892) in Creţeşti 1, and a phocid from Măluşteni. This review aims to aid the understanding of the Neogene ecosystems in the north of the Paratethys in terms of faunal contents, biostratigraphy and palaeoecology.
References
Abazzi L., Delfino M., Gallai G., Trebini L. & Rook L. (2008) – New data on the vertebrate assemblage of Fiume Santo (North-West Sardinia, Italy) and overview on the Late Miocene Tusco-Sardinian Palaeobioprovince. Palaeontology, 51(2): 425–451.
Abella J. (2011) – Indarctos arctoides Depéret, 1895 (Carnivora, Mammalia) del yacimiento vallesiense de Batallones 3 (cuenca de Madrid). PhD Thesis, Universidad Autonoma de Madrid, Madrid.
Alcalá L. (1994) – Macromammiferos neogenos de la fosa de Alfambra-Teruel. PhD Thesis, Museo Nacional de Ciencias Naturales, Teruel.
Astre G. (1929) – Sur un félin a particularités ursoïdes des limons Pliocènes de Roussillon. Bulletin de la Société Géologique de France, 29: 199–204.
Athanasiu S. (1912) – Asupra unor resturi de mamifere fosile Pliocene şi cuaternare din România. Dări de Seamă ale Institutului Geologic al României, 3: 4 pp.
Athanasiu S. (1915) – Resturile de mamifere Pliocene dela Măluşteni în Districtul Covurlui. Anuarul Institutului Geologic al României, 6: 397–408.
Audubon J.J. & Bachman J. (1851) – The Quadrupeds of North America, Vol. II. V.G. Audubon, New York, 334 pp.
Averianov A.O. & Baryshnikov G.F. (1999) – Dinofelis sp. (Carnivora, Felidae) from the Late Pliocene locality Etulia in Moldavia. Neues Jahrbuch für Geologie und Paläontologies, Monatshefte, 1999: 531–540.
Bartolini-Lucenti S. (2017) – Nyctereutes megamastoides (Canidae, Mammalia) from the early and middle Villafranchian (late Pliocene and early Pleistocene) of the lower Valdarno (Firenze and Pisa, Tuscany, Italy). Rivista Italiana di Paleontologie e Stratigrafia, 123(2): 211–218.
Bartolini-Lucenti S. (2018) – Revising the species “Mustela” ardea Gervais, 1848–1852 (Mammalia, Mustelidae): Martellictis gen. nov. and the systematics of the fossil “Galictinae” of Eurasia. Comptes Rendus Palevol, 17: 522–535.
Bartolini-Lucenti S. (2019) – “Measure my teeth and you’ll know what I ate”: The molar ratio method and an updated interpretation of the diet of Nyctereutes sinensis (Carnivora, Canidae). Acta Zoologica, 100(2): 211–215.
Bartolini-Lucenti S. & Madurell-Malapeira J. (2021) – Ancestral foxes at the gates of Europe: the Pliocene fox from Çalta-1 (Turkey) and their relationships with Asian and European Plio-Pleistocene foxes. Comptes Rendus Palevol, 20(29): 619–626.
Bartolini-Lucenti S. & Rook L. (2021) – “Canis” ferox Revisited: Diet Ecomorphology of Some Long Gone (Late Miocene and Pliocene) Fossil Dogs. Journal of Mammalian Evolution, 28: 285–306.
Bartolini-Lucenti S., Rook L. & Morales J. (2018) – Nyctereutes (Mammalia, Carnivora, Canidae) from Layna and the Eurasian Racoon-Dogs: an Updated Revision. Rivista Italiana di Paleontologia e Stratigrafia, 124(3): 597–616.
Bartolini-Lucenti S., Madurell-Malapeira J. & Rook L. (2022) – The carnivorans from Cava Monticino (Faenza, Italy; Messinian) revisited. Historical Biology, 34(8): 1458–1470.
Baryshnikov G.F. (2009) – Pleistocene Mustelidae (Carnivora) from Paleolithic site in Kudaro Caves in the Caucasus. Russian Journal of Theriology, 8(2): 75–95.
Baryshnikov G.F. & Puzachenko A.Y. (2017) – Morphometric analysis of metacarpal and metatarsal bones of cave bears (Carnivora, Ursidae). Fossil Imprint, 73(1–2): 7–47.
Batsch A.J.G.C. (1788) – Versuch einer Anleitung, zur Kenntniβ und Geschichte der Thiere und Mineralien, für akademische Vorlesungen entworfen und mit den nöthigsten Abbildungen versehen. Academischen Buchhandlung, Jena, 690 pp.
Berta A., Kienle K., Bianucci G. & Sorbi S. (2015) – A Reevaluation of Pliophoca etrusca (Pinnipedia, Phocidae) from the Pliocene of Italy: Phylogenetic and Biogeographic Implications. Journal of Vertebrate Paleontology, 35(1): e889144.
Blasius A.W.H. (1884) – Der japanische Nör, Foetorius Itatsi. Bericht der Naturforsch Gesellschaftsgründung, 13: 1–34.
Böhme M., Spassov N., Ebner M., Geraads D., Hristova L., Kirscher U., Kötter S., Linneman U., Prieto J., Roussiakis S., Theodorou G., Uhlig G. & Winklhofer M. (2017) – Messinian age and savannah environment of the possible hominin Graecopithecus from Europe. PLoS ONE, 12(5): e0177347.
Bonaparte C.L. (1838) – Synopsis vertebratorum systematis. Nuovi Annali delle Scienze Naturali, 2: 105–133.
Bonaparte C.L. (1845) – Catalogo Methodico dei Mammiferi Europei. L. di Giacomo Pirola, Milan, 36 pp.
de Bonis L. (2005) – Carnivora from the late Miocene of Akkaşdağı, Turkey. In: Sen S. (Ed.) – Geology, mammals and environments at Akkaşdağı, late Miocene of Central Anatolia. Geodiversitas, 27(4): 567–589.
de Bonis L. & Koufos G. (1991) – The late Miocene small carnivores of the lower Axios Valley (Macedonia – Greece). Geobios, 24(2): 361–379.
de Bonis L., Peigné S., Likius A., Mckaye H.T., Vignaud P. & Brunet M. (2007) – The oldest African fox (Vulpes riffautae n. sp., Canidae, Carnivora) recovered in the late Miocene deposits of the Djurab desert, Chad. Naturwissenschaften, 94: 575–580.
Bordeianu M., Valenciano A. & Codrea V.A. (2022) – Preliminary data on a medium sized carnivore from Fălciu. Simpozionul Naţional “PETRODAVA”, ediţia a III-a Piatra-Neamţ, Romania.
Boscaini A., Alba D.M., Beltrán J. F., Moyà-Solà S. & Madurell-Malapeira J. (2016) – Latest Early Pleistocene remains of Lynx pardinus (Carnivora, Felidae) from the Iberian Peninsula: taxonomy and evolutionary implications. Quaternary Science Reviews, 143: 96–106.
Bowdich T.E. (1821) – An analysis of the natural classifications of Mammalia, for the use of students and travellers. J. Smith, Paris, 115 pp.
Brisson M.J. (1762) – Le regnum animale in classes IX distributum, sive synopsis methodica sistens generalem animalium distributionem in classes IX, & duarum primarum classium, quadrupedum scilicet & cetaceorum, particularem dibvisionem in ordines, sections, genera & species. T. Haak, Paris, 296 pp.
Brünnich M.T. (1772) – Zoologiae fundamenta praelectionibus academicis accomodata. Grunde i dyrelaeren, Leipzig, 253 pp.
Del Campana D. (1914) – La Lycyaena lunensis n. sp. dell’ossario pliocenico di Olivola (Val di Magra). Paleontographia Italica, 20: 87–104.
Cherin M., Iurino D., Sardella R. & Rook L. (2014) – Acinonyx pardinensis (Carnivora, Felidae) from the Early Pleistocene of Pantalla (Italy): predatory behavior and ecological role of the giant Plio-Pleistocene cheetah. Quaternary Science Reviews, 87: 82–97.
Cipullo A. (2010) – L’evoluzione del genere Lynx dal Pliocene al Pleistocene medio nell’area Mediterranea. PhD Thesis, University of Modena-Bologna-Roma.
Coca-Ortega C. & Pérez-Claros J.A. (2019) – Characterizing ecomorphological patterns in hyenids: a multivariate approach using postcanine dentition. PeerJ, 6: e6238.
Codrea V. (1996) – Miocene rhinoceroses from Romania: an overview. Acta Zoologica Cracoviensia, 39(1): 83–88.
Crespo V., Vasile Ş., Petculescu A., Răţoi B. & Haiduc B. (2023) – The Early Pliocene small mammals (Eulipotyphla, Rodentia, Lagomorpha) from Bereşti and Măluşteni (eastern Romania): a fresh look at old collections. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 114(1–2): 49–63.
Croizet J. B. & Jobert A. (1828) – Recherches sur les ossemens fossils du Département du Puy-de-Dôme. Thibaut-Landriot, Paris, 224 pp.
Crusafont-Pairo M. & Golpe J.-M. (1962a) – Présence dans le Miocène d’Espagne de Mustélidés connus à l’est sur les bords de la Tethys. Comptes Rendus de l’Académie des Sciences Paris, 254: 2807–2809.
Crusafont-Pairó M. & Golpe-Posse J.-M. (1962b) – Nuevos hallazgos de lútridos aonicoides (Sivaonyx, Enhydriodon) en el Pikermiense Español (Cuenca de Teruel). Notas y Comunicaciones del Instituto Geológico y Minero de España, 67: 5–16.
Cuccu, A., Valenciano A., Azanza B. & DeMiguel D. (2023) – A new lynx mandible from the Early Pleistocene of Spain (La Puebla de Valverde, Teruel) and a taxonomical multivariate approach of medium-sized felids. Historical Biology, 35(1): 127–138.
Cuvier G. (1817) – Le règne animal distribué d’après son organization : pour compar de base a l’histoire naturelle des animaux et d’introduction a l’anatomie compare, t. I. Chez Déterville, Paris, 540 pp.
Cuvier G. (1824) – Recherches sur les ossemens fossils, où l’on rétablit les charatères de plusieurs animaux don’t les revolutions du globe ont détruit les espèces. d’Ocagne, Paris, 547 pp.
Daguenet T. & Sen S. (2019) – Phylogenetic relationships of Nyctereutes Temminck, 1838 (Canidae, Carnivora, Mammalia) from early Pliocene of Çalta, Turkey. Geodiversitas, 41(18): 663–677.
Depéret C. (1890) – Les Animaux Pliocènes du Roussillon. Mémoires de la Société géologique de France, 3: 5–195.
Depéret C. (1892) – La faune des mammiferes miocenes de La Crive St-Alban. Archives du Museum, Lyon, 5: 1–89.
Depéret C. (1895) – Résultats des fouilles paléontologiques dans le Miocène supérieur de la colline de Montredon. Comptes rendus hebdomadaires des Séances de l’Académie des Sciences, 121: 432–434.
Dewaele L., Lambert O. & Louwye S. (2018) – A late surviving Pliocene seal from high latitudes of the North Atlantic realm: the latest monachine seal on the southern margin of the North Sea. PeerJ, 6: e5734.
Falconer H. (1868) – Palaeontological memoirs and notes of the late Hugh Falconer, with a biographical sketch of the author. Vol. I. Fauna Antiqua Sivalensis. Robert Hardwicke, London, 590 pp.
Fejfar O. & Sabol M. (2004) – Pliocene Carnivores (Carnivora, Mammalia) from Ivanovce and Hajnáka (Slovakia). Courier Forschungsinstitut Senckenberg, 246: 15–53.
Ficarelli G. & Torre D. (1975) – Felis christoli Gervais delle sabbie plioceniche di Montpellier. Bollettino della Società Paleontologica Italiana, 14: 217–220.
Fischer von Waldheim G. (1817) – Adversaria Zoologica, fasciculus primus. Mémoires de la Société des Naturalistes de Moscou, 5: 357–446.
Forsyth Major C.I. (1901). – On Enhydrictis galictoides, a new fossil from Sardinia. Proceedings of the Zoological Society of London, 2: 625–628.
Forsyth Major C.I. (1903) – New Carnivora from the middle Miocene of La Grive St-Alban. Geological Magazine N.S., 1: 534–538.
Fraile S. (2017) - Protictitherium crassum, la pequeña hiena de 9 millones de años del Cerro de los Batallones. In: Morales J. & Baquedano E. (Eds) – La colina de los Tigres Dientes de Sable: Los yacimientos miocenos del Cerro de los Batallones (Torrejón de Velasco, Comunidad de Madrid): 221–234. Community of Madrid, Madrid.
Fraile S., Pérez B., DeMiguel D. & Morales J. (1997) – Revisión de los Carnívoros presentes en los yacimientos del Neógeno español. In: Calvo J. P. & Morales J. (Eds) – Avances en el conocimiento del Terciario Ibérico: 77–80. Universidad Complutense de Madrid, Madrid.
Frisch J.L. (1775) – Das Natur-System der vierfüssigen Thiere. C.F. Günther, Glogow, 30 pp.
Gabunia L. & Vekua A. (1998) – The find of Dinofelis in the Pliocene of Georgia. Bulletin of the Georgian Academy of Sciences, 157(2): 335–338.
García-Martínez D., Valenciano A., Suárez Bilbao A., Palancar C.A., Megía García I., Moreno D., Campaña I. & Moya-Maleno P.R. (2020) – New remains of a primitive badger from Cueva de los Toriles (Carrizosa, Castilla-La Mancha, Iberian Peninsula) suggest a new quaternary locality in the southern Iberian plateau. Journal of Iberian Geology, 46: 209–222.
Gaudry A. (1861) – Note sur les carnassiers fossiles de Pikermi (Grèce). Bulletin de la Société géologique du France (série 2), 18: 527–537.
Geraads D., Alemseged Z., Bobe R. and Reed D. (2011) – Enhydriodon dikikae, sp. nov. (Carnivora: Mammalia), a gigantic otter from the Pliocene of Dikika, Lower Awash, Ethiopia. Journal of Vertebrate Paleontology, 31: 447–453.
Geraads D. & Spassov N. (2016) – Musteloid carnivores from the upper Miocene of South-Western Bulgaria, and the phylogeny of the Mephitidae. Geodiversitas, 38(4): 543–558.
Gervais M.P. (1846) – Observations sur diverses espèces de mamifères fossiles du midi de la France. Annales des Sciences Naturelles, ser. 3, 5: 248–265.
Gervais M.P. (1848–1852) – Zoologie et Paléontologie Françaises. Nouvelles recherches sur les animaux vivants et fossiles de la France. 1st Edition. Arthus Bertrand, Paris, 271 pp.
Gervais M.P. (1859) – Zoologie et paléontologie françaises (Animaux Vertébres) ou nouvelles recherches sur les animaux vivants et fossiles de la France. 2nd Edition. Arthus Bertrand, Paris, 544 pp.
Ghenea C. (1968) – Studiul depozitelor Pliocene dintre Valea Prutului şi Valea Bîrladului. Comitetul de Stat al Geologiei, Institutul Geologic, Studii Tehnice şi Economice, Seria J, Stratigrafie, 6: 1–137.
Ginsburg L. (1998) – Le gisement de vertébrés pliocènes de Çalta, Ankara, Turquie. 5. Carnivores. Geodiversitas, 20: 379–396.
Gray J.E. (1821) – On the natural arrangement of vertebrose animals. London Medical Repository, 15: 296–310.
Gray J.E. (1825) – An outline of an attempt at the disposition of the Mammalia into tribes and families with a list of the genera apparently appertaining to each tribe. Annals of Philosophy (n. s.), 10: 337–344.
Gray J.E. (1844–1875) – The zoology of the voyage of H.M.S. Erebus and Terror, under the command of Captain Sir James Clark Ross, N.N., F.R.S., during the years 1839 to 1843. Vol. 1. Mammalia, Birds. E.W. Janson, London, 53 pp.
Gray J.E. (1865) – Revision of the genera and species of Mustelidae contained in the British Museum. Proceedings of the Geological Society of London, 1865: 100–154.
Grohé C., de Bonis L., Chaimanee Y., Blondel C. & Jaeger J.-J. (2013) – The oldest Asian Sivaonyx (Lutrinae, Mustelidae): a contribution to the evolutionary history of bunodont otters. Palaeontologia Electronica, 16.3.29A: 1–13.
Grohé C., de Bonis L., Chaimanee Y., Chavasseau O., Rugbumrung M., Yamee C., Suraprasit K., Gibert C., Surault J., Blondel C. & Jaeger J.-J. (2020) – The late middle Miocene Mae Moh Basin of northern Thailand: the richest Neogene assemblage of Carnivora from Southeast Asia and a paleobiogeographic analysis of Miocene Asian carnivorans. American Museum Novitates, 3952: 1–57.
Grohé C., Uno K. & Boisserie J.-R. (2022) – Lutrinae Bonaparte, 1838 (Carnivora, Mustelidae) from the Plio-Pleistocene of the Lower Omo Valley, southwestern Ethiopia: systematics and new insights into the paleoecology and paleobiogeography of the Turkana otters. Comptes Rendus Palevol, 21(30): 681–705.
Hammer Ø., Harper D.A.T. & Ryan P.D. (2001) – PAST: Paleontological Statistics Software Package for Education and Data Analysis. Palaeontologia Electronica, 4(1): 1–9.
Hay O.P. (1921) – Descriptions of species of Pleistocene Vertebrata, types or specimens most of which are preserved in the United States National Museum. Proceedings of the United States National Museum, 59: 599–642.
Howell F.C. & Petter G. (1980) – The Pachycrocuta and Hyaena lineages (Plio-Pleistocene and extant species of the Hyaenidae). Their Relationships with Miocene ictitheres: Palhyaena and Hyaenictitherium. Geobios, 13(4): 579–623.
Howell F.C. & Petter G. (1985) – Comparative observations on some Middle and Upper Miocene hyaenids. Genera: Percrocuta Kretzoi, Allohyaena Kretzoi, Adcrocuta Kretzoi (Mammalia, Carnivora, Hyaenidae). Geobios, 18(4): 419–476.
Hugueney M. (1975) – Les mustelids (Mammalia, Carnivora) du gisement pleistocene moyen de la Fage (Correze). Nouvelles archives du Muséum d’histoire naturelle de Lyon, 13: 29–46.
Hürzeler J. (1987) – Die Lutrinen (Carnivora, Mammalia) aus dem «Grosseto-Lignit» der Toscana. Schweizerische Paläontologische Abhandlungen, 110: 28–48.
Hürzeler J. & Engesser B. (1976) – Les faunes de mammifères néogènes du Bassin de Baccinello (Grosseto, Italie). Comptes Rendus de l’Académie des Sciences, Paris, 283: 333–336.
Ionesi L. (1994) – Geologia unităţilor de Platformă şi a Orogenului Nord-Dobrogean. Bucharest, Editura Tehnică, 280 pp.
Jeanrenaud P. (1971) – Geologia Moldovei Centrale dintre Siret şi Prut. PhD Thesis, Alexandru Ioan Cuza University of Iaşi.
Karadenizli L., Seyitoğlu G., Sen S., Arnaud N., Kazanci N., Saraç G. & Alçiçek C. (2005) – Mammal bearing late Miocene tuffs of the Akkaşdağı region; distribution, age, petrographical and geochemical characteristics. In: Sen S. (Ed.) – Geology, mammals and environments at Akkaşdağı, late Miocene of Central Anatolia. Geodiversitas, 27(4): 553–566.
Kampouridis P., Răţoi B. & Ursachi L. (2023) – New evidence for the unique coexistence of two subfamilies of clawed perissodactyles (Mammalia, Chalicotheriidae) in the Upper Miocene of Romania and the Eastern Mediterranean. Journal of Mammalian Evolution, 30: 641–656.
Kargopoulos N. (2019) – Carnivoran dietary adaptations: a multiproxy study on the feeding ecology of the fossil carnivorans of Greece. MSc Thesis, National and Kapodistrian University of Athens.
Kargopoulos N., Kampouridis P., Lechner T. & Böhme M. (2021a) – A review of Semigenetta (Viverridae, Carnivora) from the Miocene of Eurasia based on material from the hominid locality of Hammerschmiede (Germany). Geobios, 69: 25–36.
Kargopoulos N., Roussiakis S., Kampouridis P. & Koufos G. (2023) – Interspecific competition in ictitheres (Carnivora: Hyaenidae) from the Late Miocene of Eurasia. Comptes Rendus Palevol, 22(3): 33–44.
Kargopoulos N., Valenciano A., Abella J., Kampouridis P., Lechner T. & Böhme M. (2022) – The exceptionally high diversity of small carnivorans from the Late Miocene hominid locality of Hammerschmiede (Bavaria, Germany). PLoS ONE, 17(7): e0268968.
Kargopoulos N., Valenciano A., Kampouridis P., Lechner T. & Böhme M. (2021b) – New species of Vishnuonyx (Carnivora, Lutrinae) from the early Late Miocene hominid locality of Hammerschmiede, Bavaria, Germany. Journal of Vertebrate Paleontology, 41(3): e1948858.
Kaya T., Geraads D. & Tuna V. (2005) – A new Late Miocene mammalian fauna in the Karaburun Peninsula (W Turkey). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 236(3): 321–349.
Khomenko I.P. (1914) – Meoticheskaya fauna s. Taraklii benderskogo uezda: Fissipedia, Rodentia, Rhinocerotinae, Equinae, Suidae, Proboscidae. Trudy Bessarahskoe Obshchestvo Estestvoispytatelei, Kishinev, 5: 1–55.
Khomenko I.P. (1932) – Hyaena borissiaki n. sp. iz russil’onskoj fauny Bessarabii. Travaux de l’Institut Paleozoologique de l’Académie des Sciences de l’U.R.S.S., 1: 81–134.
Koretsky I. (2001) – Morphology and Systematics of Miocene Phocinae (Mammalia: Carnivora) from Paratethys and the North Atlantic Region. Geologica Hungarica, 54: 1–109.
Koretsky I. & Holec P. (2002) – A Primitive Seal (Mammalia: Phocidae) from the Early Middle Miocene of Central Paratethys. Smithsonian Contributions to Paleobiology, 93: 163–178.
Koretsky I. & Rahmat S. (2013) – First record of fossil Cystophorinae (Carnivora, Phocidae): Middle Miocene seals from the Northern Paratethys. Rivista Italiana di Paleontologia e Stratigrafia, 119(3): 325–350.
Koretsky I. & Rahmat S. (2015) – A new species of the subfamily Devinophocinae (Carnivora, Phocidae) from the Central Paratethys. Rivista Italiana di Paleontologia e Stratigrafia, 121(1): 31–47.
Koretsky I., Peters N. & Rahmat S. (2015) – New species of Praepusa (Carnivora, Phocidae, Phocinae) from the Netherlands supports East to West Neogene dispersal of true seals. Vestnik zoologii, 49(1): 57–66.
Kormos T. (1932) – Neue pliozäne Nagetiere aus der Moldau. Paläontologische Zeitschrift, 14: 193–199.
Kormos T. (1934) – Neue und wenig bekannte Musteliden aus dem ungarischen Oberpliozän. Folia Zoologica et Hydrobiologica, 5(2): 129–157.
Kostopoulos D., Koufos G., Sylvestrou I., Syrides G. & Tsombachidou E. (2009) – The Late Miocene Mammal Faunas of the Mytilinii Basin, Samos Island, Greece: New Collection. 2. Lithostratigraphy and Fossiliferous Sites. Beiträge zur Paläontologie, 31: 13–26.
Koufos G. (1997) – The canids Eucyon and Nyctereutes from the Ruscinian of Macedonia, Greece. Paleontologia i Evolució, 30/31: 39–48.
Koufos G. (2006) – The Neogene mammal localities of Greece: Faunas, chronology and biostratigraphy. Hellenic Journal of Geosciences, 41: 183–214.
Koufos G. (2009) – The Late Miocene Mammal Faunas of the Mytilinii Basin, Samos Island, Greece: New Collection. 5. Carnivora. Beiträge zur Paläontologie, 31: 57–105.
Koufos G. (2016) – Palaeontology of the upper Miocene vertebrate localities of Nikiti (Chalkidiki Peninsula, Macedonia, Greece). Carnivora. Geobios, 49: 53–67.
Koufos G. (2021) – The late Miocene middle-sized ictitheres (Mammalia: Hyaenidae) of Greece. Bollettino della Società Paleontologica Italiana, 60(1): 19–48.
Kowalski, K. (1970) – Les Mammiferes Fossiles des Remplissages Karstiques de Pologne—Problemes Paleobiologiques et Stratigraphiques. In: Orghidan T. & Racovitza G. (Eds) – Livre du Centenaire Emile G. Racovitza, 1868–1968: 491–498. Bucharest, Académie de la République Socialiste de Roumanie.
Kretzoi M. (1938) – Die Raubtiere von Gombaszög nebst einer Übersicht der Gesamtfauna. Annales Museum Nationale Hungaricum, 31: 89–157.
Kretzoi M. (1943) – Kochitis centenii n. g. n. sp., ein altertümlicher Creodonte aus dem Oberoligozän Siebenbürgens. Földtani Közlöny, 73: 10–195.
Kretzoi M. (1945) – Bemerkungen über das Raubtiersystem. Annales Musei historico-naturalis hungarici, 38: 59–83.
Kurtén B. (1978) – The lynx from Etouaires, Lynx issiodorensis (Croizet & Jobert), late Pliocene. Annales Zoologici Fennici, 15: 314–322.
Kurtén B. (1982) – Status of the Fossil Hyaenids Ictitherium viverrinum and Thalassictis robusta (Mammalia). Zeitschrift für Geologische Wissenschaften, 10(7): 1009–1018.
Kurtén B. & Werdelin L. (1984) – The relationships of Lynx shansius Teilhard. Annales Zoologici Fennici, 21: 129–133.
Kurtén B. & Werdelin L. (1988) – A review of the genus Chasmaporthetes Hay, 1921 (Carnivora, Hyaenidae). Journal of Vertebrate Paleontology, 8(1): 46–66.
Lartet E. (1851) – Notice sur la colline de Sansan, suivie d’une récapitulation des diverses espèces d’animaux vertébrés fossiles, trouvés soit à Sansan, soit dans d’autres gisements du terrain tertiaire miocène dans le bassin souspyrénéen. J.A. Portes, Auch, 45 pp.
Lesson R.P. (1827) – Manuel de mammalogie, ou histoire naturelle des mammiferes. J.B. Bailliere, Paris, 441 pp.
Linnaeus C. (1758) – Systema naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Laurentius Salvius, Stockholm, 823 pp.
Linnaeus C. (1767) – Systema naturae per regna tria naturae editio duodecima reformata 1 (Regnum animale). Laurentius Salvius, Stockholm, 1347 pp.
Lungu A & Rzebik-Kowalska B. (2011) – Faunal assemblages, stratigraphy and taphonomy of the Late Miocene localities in the Republic of Moldova. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 62 pp.
Macarovici N. (1957) – Mammifères fossils du Sarmatien de Păun (Jassy). Analele Ştiinţifice, 2(4): 143–154.
Macarovici N. & Paghida N. (1966) – Flora şi fauna din Sarmaţianul dela Păun-Iaşi. Analele Universităţii Bucureşti, Ştiinţele Naturii, Geologie Geografie, 15 (1): 67–81.
Marciszak A. (2016) – The common weasel Mustela nivalis L., 1766 from Sarakenos Cave (Greece) and the difficulty to distinguish small mustelid species in the fossil record. Acta Zoologica Cracoviensia, 59(1): 25–35.
Marciszak A., Kropczyk A., Gornig W., Kot M., Nadachowski A. & Lipecki G. (2023) – History of Polish Canidae (Carnivora, Mammalia) and Their Biochronological Implications on the Eurasian Background. Genes, 14: 539.
Marciszak A., Lipecki G. & Spassov N. (2024) – Martes wenzensis Stach, 1959 within the early history of the genus Martes Pinel, 1792. Acta Geologica Polonica, 74(1): e4.
Martin R. (1973) – Trois nouvelles espèces de Caninae (Canidae, Carnivora) des gisements Plio-villafranchiens d’Europe. Documents du Laboratoire de Géologie de la Faculté des Sciences de Lyon, 57: 87–96.
Mecozzi B., Sardella R., Boscaini A., Cherin M., Costeur L., Madurell-Malapeira J., Pavia M., Profico A. & Iurino D. (2021) – The tale of a short-tailed cat: New outstanding Late Pleistocene fossils of Lynx pardinus from southern Italy. Quaternary Science Reviews, 262: 106840.
Meneghini G. (1863) – Descrizione dei Resti di due Fiere: Trovati nelle Ligniti Mioceniche di Montebamboli. Atti della Società italiana di scienze naturali, 4: 17–33.
Monguillon A., Spassov N., Argant A., Kauhala K. & Viranta S. (2004) – Nyctereutes vulpinus comb. et stat. nov. (Mammalia, Carnivora, Canidae) du Pliocène terminal de Saint-Vallier (Drôme, France). Geobios, 37: 83–88.
Montoya P., Ginsburg L., Alberdi M. T., Van der Made J., Morales J. & Soria M.D. (2006) – Fossil large mammals from the early Pliocene locality of Alcoy (Spain) and their importance in biostratigraphy. Geodiversitas, 28(1): 137–173.
Montoya P., Morales J. & Abella J. (2009) – Eucyon debonisi n. sp., a new Canidae (Mammalia, Carnivora) from the latest Miocene of Venta del Moro (Valencia, Spain). Geodiversitas, 31(4): 709–722.
Montoya P., Morales J. & Abella J. (2011) – Musteloidea (Carnivora, Mammalia) from the late Miocene of Venta del Moro (Valencia, Spain). Estudios Geológicos, 67: 193–206.
Morales J. (1984) – Venta del Moro: su macrofauna de mamiferos, y biostratigrafia continental del Miocene terminal Mediterraneo. PhD Thesis, Universidad Complutense de Madrid.
Morales J. (2016) – Los carnìvoros de Villarroya. In: Alberdi M.T., Azanza B. & Cervantes E. (Eds) – Villaroya, Yacimiento clave de la paleontologia Riojana: 119–142. Instituto de Estudios Riojanos, Logroño.
Morales J. & Pickford M. (2005) – Giant Bunodont Lutrinae from the Mio-Pliocene of Kenya and Uganda. Estudios Geológicos, 61: 233–246.
Morales J., Soria D., Montoya P., Pérez B. & Salesa M.J. (2003) – Caracal depereti nov. sp. y Felis aff. silvestris (Felidae, Mammalia) del Pliocene inferior de Layna (Soria, España). Estudios Geológicos, 59: 229–247.
Morales J., Pickford M. & Soria D. (2005) – Carnivores from the late Miocene and basal Pliocene of the Tugen Hills, Kenya. Revista de la Sociedad Geológica de España, 18: 39–61.
Morlo M. & Kundrát M. (2001) – The first carnivoran fauna from the Ruscinium (Early Pliocene, MN 15) of Germany. Paläontologische Zeitschrift, 75(2): 163–187.
de Muizon C.D. & Hendey Q.B. (1980) – Late Tertiary seals of the South Atlantic Ocean. Annals of the South African Museum, 2: 91–128.
Nadachowski A., Mirosław-Grabowska J., David A., Tomek T., Garapick A., Pascaru V., Obadă T. & Szyndlar Z. (2006) – Faunal assemblages and biostratigraphy of several Pliocene sites from Moldova. Courier Forschungsinstitut Senckenberg, 256: 249–259.
Nagel D., Pronin K., Pytlik I., Urbanek C., Ivanoff D. & Semenov Y. (2004) – Die Pliozäne Fauna der Katakomben von Odessa (Ukraine). Berichte des Institutes für Erdwissenschaften der Karl-Franzens-Universität Graz, 9: 278–280.
Newton E.T. (1890) – On some New Mammals from the Red and Norwich Crags. The Quarterly Journal of the Geological Society of London, 46: 444–453.
Newton E.T. (1894) – The Vertebrate Fauna collected by Mr. Lewis Abbott from the Fissure near Ightham, Keny. The Quarterly Journal of the Geological Society of London, 50: 188–211.
Odintsov I.A. (1965) – Vulpes praecorsac Kormos from Pliocene deposits of Odessa. Paleontologichesky Sbornik, Lvov University, 2: 57–64.
Odintsov I.A. (1967) – New species of Pliocene Carnivora, Vulpes odessana sp. nov. from the Karst Cave of Odessa. Paleontologichesky Sbornik, Lvov University, 4: 130–137.
Palcu D.V., Golovina L.A., Vernyhorova Y.V., Popov S.V. & Krijgsman W. (2017) – Middle Miocene paleoenvironment crises in Central Eurasia caused by changes in marine gateway configuration. Global and Planetary Change, 158: 57–71.
Palcu D.V., Vasiliev I., Stoica M. & Krijgsman W. (2019) – The end of the Great Khersonian Drying of Eurasia: Magnetostratigraphic dating of the Maeotian transgression in the Eastern Paratethys. Basin Research, 31: 33–58.
Palmqvist P., Torregrosa V., Pérez-Claros J.A., Martínez-Navarro B. and Turner A. (2007) – A re-evaluation of the diversity of Megantereon (Mammalia, Carnivora, Machairodontinae) and the problem of species identification in extinct carnivores. Journal of Vertebrate Paleontology, 27(1): 160–175.
Paparizos N., Kargopoulos N., Liakopoulou D. & Roussiakis S. (Submitted) – New material of Hyaenictis graeca (Carnivora, Hyaenidae) from the Late Miocene of Greece. Journal of Vertebrate Paleontology.
Pei W.C. (1934) – On the Carnivora from locality 1 of Choukoutien. Palaeontologia Sinica, 13(1): 1–217.
Peigné S. (2016) – Carnivora. In Sen S. (Ed) – Late Miocene mammal locality of Küçükçekmece, European Turkey. Geodiversitas, 38(2): 197–224.
Pevzner M.A., Vangengeim E.A., Vislobokova I.A., Sotnikova M.V. & Tesakov A.S. (1996) – Ruscinian of the territory of the former Soviet Union. Newsletters on Stratigraphy, 33(2): 77–97.
Philipsson L.M. (1788) – Dissertatio historiconaturalis sistens nova testaceorum genera. Typis Berlingianis, Lund, 23 pp.
Pickford M. (2007) – Revision of the Mio-Pliocene bunodont otter-like mammals of the Indian Subcontinent. Estudios Geológicos, 63(10): 83–127.
Pilgrim G.E. (1931) – Catalogue of the Pontian Carnivora of Europe in the Department of Geology. British Museum of Natural History, London, 174 pp.
Pilgrim G.E. (1932) – The fossil Carnivora of India. Memoirs of the Geological Survey of India. Palaeontologia Indica, 18: 1–232.
Pilgrim G.E. (1933) – A fossil skunk from Samos. American Museum Novitates, 663: 1–15.
Pinel P. (1792) – Recherches sur une nouvelle classification des quadrupes, fondee sur la structure mehanique des parties osseuses qui servent al’ articulation de la machoire inferieure. Actes de la société d’histoire naturelle de Paris, 1: 50–60.
Pocock R.I. (1921) – On the External Characters and Classification of the Mustelidae. Proceedings of the Zoological Society of London, 1921: 803–837.
Pomel M. (1843) – Nouvelle espèce de chien fossile découverte dans les alluvions volcaniques de l’Auvergne. Bulletin de la Société Géologique de France, 14: 38–41.
Qiu Z.-X. (1987) – Die Hyaeniden aus dem Ruscinium und Villafranchium Chinas. Münchner Geowissenschaftliche Abhandlungen, Reihe A: Geologie und Palaontologie, 9: 1–110.
Radulescu C. & Kisgyörgy Z. (1970) – Contribution a la connaissance de la faune de mammiferes villafranchiens de Căpeni-Vîrghis (Dépression de Braşov). Aluta, 1970: 11–23.
Radulescu C., Samson P.-M., Petculescu A. & Stiucã E. (2003) – Pliocene Large Mammals of Romania. Coloquios de Palaeontología, Vol. Ext. 1: 549–558.
Rahmat S., Muniz F., Toscano A., Esperante R. & Koretsky I. (2020) – First European record of Homiphoca (Phocidae: Monachinae: Lobodontini) and its bearing on the paleobiogeography of the genus. Historical Biology, 32(4): 561–569.
Răţoi B.G. & Brânzilă M. (2012) – Sedimentological study of the Khersonian deposits from Paiu and Monument open pits (Vaslui City): Preliminary data. Studii şi Cercetări Geology and Geography, Bistriţa, 17: 71–80.
Reynolds S.A. (1912) – Monograph of the British Pleistocene Mammalia. Vol II. British Pleistocene Hyaenidae, Ursidae, Canidae and Mustelidae. Part IV. The Mustelidae. The Palaeontographical Society, London, 28 pp.
Rivals F., Belyaev R.I., Basova V.B. & Prilepskaya N. (2024) – A tale from the Neogene savanna: Paleoecology of the hipparion fauna in the northern Black Sea region during the Late Miocene. Palaeogeography, Palaeoclimatology, Palaeoecology, DOI:10.1016/j.palaeo.2024.112133.
Rook L. (1993) – I Cani dell’Eurasia dal Miocene superiore al Pleistocene medio. PhD Thesis, Universitá di Firenze.
Rook L., Bartolini-Lucenti S., Bukhsianidze M. & Lordkipanidze D. (2017) – The Kvabebi Canidae record revisited (late Pliocene, Sighnaghi, eastern Georgia). Journal of Paleontology, 91(6): 1258–1271.
Rook L., Ficcarelli G. & Torre D. (1991) – Messinian carnivores from Italy. Bollettino della Società Paleontologica Italiana, 30: 7–22.
Rook L., Gallai G. & Torre D. (2006) – Lands and endemic mammals in the Late Miocene of Italy: Constrains for paleogeographic outlines of Tyrrhenian area. Palaeogeography, Palaeoclimatology, Palaeoecology, 238(1–4): 263–269.
Rook L., Bartolini Lucenti S., Tuveri C. & Arca M. (2018) – Mustelids (Carnivora, Mammalia) from Monte Tuttavista fissure fillings (Early and Middle Pleistocene; Orosei, Sardinia): Taxonomy and evolution of the insular Sardinian Galictini. Quaternary Science Reviews, 197: 209–223.
Roth J. & Wagner A. (1854) – Die fossilen Knochenüberreste von Pikermi in Griechenland. Abhandlungen der Bayerisehen Akademie der Wissensehaften, 7: 371–464.
Salesa M., Antón M., Morales J. & Peigne S. (2012a) – Systematics and phylogeny of the small felines (Carnivora, Felidae) from the Late Miocene of Europe: a new species of Felinae from the Vallesian of Batallones (MN 10, Madrid, Spain). Journal of Systematic Palaeontology, 10(1): 87–102.
Salesa M., Pesquero M.D., Siliceo G., Antón M., Alcalá L. & Morales J. (2012b) – A rich community of Felidae (Mammalia, Carnivora) from the late Miocene (Turolian, MN 13) site of Las Casiones (Villalba Baja, Teruel, Spain). Journal of Vertebrate Paleontology, 32(2): 658–676.
Samson P. & Radulesco C. (1973) – Les faunes de mammifères et la limite Pliocène–Pléistocène en Roumanie. Travaux de l’Institut de Spéologie “Émile Racovitza”, 12: 191–228.
Săndulescu M. (1984) – Geotectonica României. Bucharest, Editura Tehnică, 335 pp.
Sato J.J., Hosoda T., Wolsan M., Tsuchiya K., Yamamoto M. & Suzuki H. (2003) – Phylogenetic relationships and divergence times among mustelids (Mammalia: Carnivora) based on nucleotide sequences of the nuclear interphotoreceptor retinoid binding protein and mitochondrial cytochrome b genes. Zoological Science, 20: 243–264.
Schlosser M. (1899) – Parailurus anglicus und Ursus böckhi aus den Ligniten von Baróth-Köpecz, Comitat Háromszék in Ungard. Mittheilungen aus dem Jahrbuche der Königlich Ungarischen Geologischen Anstalt, 13: 66–95.
Schlosser M. (1903) – Die fossilen Säugethiere Chinas nebst einer Odontographie der recenten Antilopen. Abhandlungender Mathematisch-Physikalischen Klasse der Königlich Bayerischen Akademie der Wissenschaften, 22: 1–220.
Schmidt-Kittler N. (1995) – The Vertebrate Locality Maramena (Macedonia, Greece) at the Turolian-Ruscinian Boundary (Neogene). 7. Carnivora (Mammalia). Münchner Geowissenschaftliche Abhandlungen (A), 28: 75–86.
Semenov Y. (1989) – Ictitheres and morphologically similar hyaenas from the Neogene of the USSR. PhD Thesis, Naukova dumka, Kiev.
Semenov Y. (2001) – Stratigraphic distribution of the terrestrial carnivores in the Vallesian and Turolian of Ukraine. Beiträge zur Paläontologie, 26: 139–144.
Semenov Y. (2008) – Taxonomical reappraisal of “ictitheres” (Mammalia, Carnivora) from the Late Miocene of Kenya. Comptes Rendus Palevol, 7(8): 529–539.
Simionescu I. (1922) – Fauna vertebrată dela Măluşteni. Anuarul Institutului Geologic al României, 9: 451–458.
Simionescu I. (1930) – Vertebratele Pliocene dela Măluşteni (Covurlui). Academia Română, Publicaţiunile Fondului Vasile Adamachi, 9(49): 83–151.
Simionescu I. (1932) – Les vertébrés pliocènes de Bereşti. Buletinul Societăţii Române de Geologie, 1: 215–228.
Simionescu I. (1938) – Mamiferele pliocene de la Cimişlia. I Carnivore. Academia Română, Publicaţiunile Fondului Vasile Adamachi, 9(50): 1–30.
Soria D. & Aguirre E. (1976) – El cánido de Layna: revisión de los Nyctereutes fósiles. Trabajos Neógeno y Cuaternario, 5: 83–115.
Sotnikova M.V. (1994) – The genus Chasmaporthetes Hay, 1921 from the Pliocene of Russia, Ukraine, Mongolia and Tadzhikistan. In: Tatarinov L.P. (Ed) – Palaeotheriology: 113–139. Nauka, Moscow.
Sotnikova M.V. (2008) – Carnivora assemblages of the Ruscinian-Early Villafranchian transition: Eastern Europe (Ukraine) and Russia (Transbaikalia) – similarity and distinctions. 6th Meeting of the European Association of Vertebrate Palaeontology, Spišská Nová Ves: 85–88.
Spassov N. (2001) – Zorillas (Carnivora, Mustelidae, Ictonychini) from the Villafranchian of Bulgaria with a description of a new species of Baranogale Kormos, 1934. Geodiversitas, 23(1): 87–104.
Spassov N. & Geraads D. (2015) – A New Felid from the Late Miocene of the Balkans and the Contents of the Genus Metailurus Zdansky, 1924 (Carnivora, Felidae). Journal of Mammalian Evolution, 22(1): 45–56.
Spassov N. & Spiridonov G. (1993) – Vormela peregusna (Guldenstaedt, 1770), In: Stubbe M. & Krapp F. (Eds) – Handbuch der Säugetiere Europas: 817–855. Aula, Wiesbaden.
Stach J. (1959) – On some Mustelinae from the Pliocene bone Breccia of Węże. Acta Palaeontologica Polonica, 4(2): 101–118.
Stefaniak K., Ratajczak U., Kotowski A., Kozlowska M. & Mackiewicz P. (2020) – Polish Pliocene and Quaternary deer and their biochronological implications. Quaternary International, 546: 64–83.
Ţabără D. & Chirilă G. (2012) – Palaeoclimatic estimation from Miocene of Romania, based on palynological data. Carpathian Journal of Earth and Environmental Sciences, 7(2): 195–208.
Tavani G. (1941) –- Revisione dei resti del Pinnipede conservato nel Museo di Geologia di Pisa. Palaeontographia Italica, 40: 97–113.
Tedford R.H. & Qiu Z. (1991) – Pliocene Nyctereutes (Carnivora: Canidae) from Yushe, Shanxi, with comments on Chinese fossil raccoon-dogs. Vertebrata PalAsiatica, 29(3): 176–189.
Tedford R.H. & Qiu Z. (1996) – A new canid genus from the Pliocene of Yushe, Shanxi Province. Vertebrata PalAsiatica, 34: 27–48.
Tedford R.H., Wang X. & Taylor B.E. (2009) – Phylogenetic systematics of the North American fossil Caninae (Carnivora: Canidae). Bulletin of American Museum of Natural History, 325: 1–218.
Teilhard de Chardin P. & Leroy P. (1945) – Les Mustélidés de Chine. Publ Inst Geobiol, 12: 1–56.
Temminck C. (1838) – Over de kennis en de Verbreiding der Zoogdieren van Japan. Tijdschr. Nat. Gesch. Phys., 5: 273–293.
Terhune C.E., Curran S., Croitor R., Drăguşin V., Gaudin T., Petculescu A., Robinson C., Robu M. & Werdelin L. (2020) – Early Pleistocene fauna of the Olteţ River Valley of Romania: Biochronological and biogeographic implications. Quaternary International, 553: 14–33.
Terzea E. (1997) – Biochronologie du Pliocene du bord meridional du basin Dacique (Roumanie). Memoires et Travaux de l’EPHE, Institut de Montpellier, 21: 649–660.
Tesakov A.S., Titov V.V., Simakova A.N., Frolov P.D., Syromyatnikova E.V., Kurshakov S.V., Trikhunkov Y.I., Sotnikova M.V., Kruskop S.V., Zelenkov N.V., Tesakova E.M. & Palatov D.M. (2017) – Late Miocene (Early Turolian) vertebrate faunas and associated biotic record of the Northern Caucasus: Geology, palaeoenvironment, biochronology. Fossil Imprint, 73(3–4): 383–444.
Tong H., Zhang B., Chen X., Jiangzuo Q., Liu J. & Wang X. (2023) – New carnivoran remains from the Early Pleistocene Shanshenmiaozui site in Nihewan Basin, northern China. Quaternary International, 658: 60–79.
Topachevski V.A. (1961) – Novyi Pliotsenovykh vid vykhukholi iz predkavkazya. Paleontologicheskij Zhurnal, 4: 131–137.
Trouessart E. L. (1897) – Catalogus Mammalium tam Vivientum quam Fossilium. Berolini, R. Friedländer & Sohn, Berlin, 1469 pp.
Tseng Z.J. & Wang X. (2007) – The first record of the late Miocene Hyaenictitherium hyaenoides Zdansky (Carnivora: Hyaenidae) in Inner Mongolia and an evaluation of the genus, Journal of Vertebrate Paleontology, 27(3): 699–708.
Tseng Z.J., Li Q. & Wang X. (2013) – A new cursorial hyena from Tibet, and analysis of biostratigraphy, paleozoogeography, and dental morphology of Chasmaporthetes (Mammalia, Carnivora). Journal of Vertebrate Paleontology, 33(6): 1457–1471.
Turner A., Antón M. & Werdelin L. (2008) – Taxonomy and evolutionary patterns in the fossil Hyaenidae of Europe. Geobios, 41: 677–687.
Ursachi L. (2016) – Asociaţii de vertebrate continentale Miocene din zona centrală a podişului Bârladului. PhD Thesis, Universitatea Al. I. Cuza Iaşi and Universitatea Babeş-Bolyai Cluj-Napoca.
Ursachi L., Codrea V., Vencz M., Solomon A. & Răţoi B. (2015) – Creţeşti-Dobrina 1: a new Early Vallesian locality in Moldova. Tenth Romanian Symposium on Paleontology Abstracts and Field Tripe Guide, Cluj Napoca, p. 115.
Valenciano A. & Govender R. (2020) – New insights into the giant mustelids (Mammalia, Carnivora, Mustelidae) from Langebaanweg fossil site (West Coast Fossil Park, South Africa, early Pliocene). PeerJ, 8: e9221.
Valenciano A., Morales J., Azanza B. & DeMiguel D. (2022b) – Aragonictis araid gen. et sp. nov., a small-sized hypercarnivore (Carnivora, Mustelidae) from the late middle Miocene of the Iberian Peninsula (Spain). Journal of Vertebrate Paleontology, 41(5): e2005615.
Valenciano A., Morales J. & Govender R. (2022a) – Eucyon khoikhoi sp. nov. (Carnivora: Canidae) from Langebaanweg ‘E’ Quarry (early Pliocene, South Africa): the most complete African canini from the Mio-Pliocene. Zoological Journal of the Linnean Society, 194(2): 366–394.
Valenciano A., Pérez-Ramos A., Abella J. & Morales J. (2020) – A new hypercarnivorous mustelid (Mammalia, Carnivora, Mustelidae) from Batallones, late Miocene (MN10), Torrejón de Velasco, Madrid, Spain. Geodiversitas, 42(8): 103–121.
Van Baak C.G.C., Mandic O., Lazar I., Stoica M. & Krijgsman W. (2015) – The Slanicul de Buzau section, a unit stratotype for the Romanian stage of the Dacian Basin (Plio-Pleistocene, Eastern Paratethys). Palaeogeography, Palaeoclimatology, Palaeoecology, 440: 594–613.
Van Beneden P.J. (1876) – Les phoques fossiles du bassin d’Anvers. Bulletins de l’Académie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 41: 783–802.
van Dam J.A., Mein P., Garcés, van Balen R.T., Furió M. & Alcalá L. (2023) – A new rodent chronology for the late Neogene of Spain. Geobios, 76: 53–74.
Vangengeim E.A., Pevzner M.A. & Tesakov A.S. (2005) – Ruscinian and Lower Villafranchian: Age of Boundaries and Position in Magnetochronological Scale. Stratigraphy and Geological Correlation, 13(50): 530–546.
Villalta Comella J.F. & Crusafont-Pairó M. (1945) – Enhydriodon lluecai n. sp. el primer lútrido del pontiense Español. Boletín de la Sociedad Española de Historia Natural, 43: 383–396.
Vinuesa V., Madurell-Malapeira J., Ansón M. & Alba D. (2014) – New cranial remains of Pliocrocuta perrieri (Carnivora, Hyaenidae) from the Villafranchian of the Iberian Peninsula. Bolletino della Società Paleontologica Italiana, 53(1): 39–47.
Vinuesa V., Madurell-Malapeira J., Werdelin L., Robles J.M., Obradó P. & Alba D. (2017) – A New Skull of Hyaenictis Gaudry, 1861 (Carnivora, Hyaenidae) Shows Incipient Adaptations to Durophagy. Journal of Mammalian Evolution, 24(2): 207–219.
Wagner A. (1837) – First Part of the nineteenth Volume of Asiatic Researches; or Transactions of the Society, instituted in Bengal, for enquiring into the History, etc. Gelehrte Anzeigen, 5(170): 334–336.
Wagner A. (1848) – Urweltliche Säugthier-Ueberreste aus Griechenland. Abhandlungen der Bayerischen Akademie der Wissenschaften 5(2): 335–378.
Wagner A. (1857) - Neue Beiträge zur Kenntniss der fossilen Säugthier-Ueberreste von Pikermi. Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathematischnaturwissenschaftlichen Klasse, 8: 111–158.
Wagner J. (2010) – Pliocene to early Middle Pleistocene ursine bears in Europe: a taxonomic overview. Journal of the National Museum (Prague), Natural History Series, 179(20): 197–215.
Wagner J., Čermák S. & Horáček I. (2011) – The presence of Ursus ex gr. minimus-thibetanus in the Late Villányan and its position among the Pliocene and Pleistocene black bears in Europe. Quaternaire, Hors-série 4: 39–58.
Wagner J., Lipecki G. & Krawczyk M. (2008) – New evidence of Ursus minimus from the territory of Poland. Stalactite, 58(2): 78–80.
Wang D., Xing L., Rothschild B., Persons S. IV & Liu J. (2023) – A palaeopathological specimen of the Late Miocene Parataxidea sp. (Mammalia: Carnivora) from the Linxia Basin, China. Historical Biology, 35(7): 1255–1260.
Wang X. & Qiu Z.-X. (2004) – Late Miocene Promephitis (Carnivora, Mephitidae) from China. Journal of Vertebrate Paleontology, 24: 721–731.
Werdelin L. (1981) – The evolution of lynxes. Annales Zoologici Fennici, 18: 37–71.
Werdelin L. (1988) – Studies of fossil hyaenas: the genera Thalassictis Gervais ex Nordmann, Palhyaena Gervais, Hyaenictitherium Kretzoi, Lycyaena Hensel and Palinhyaena Qiu, Huang & Guo. Zoological Journal of the Linnaean Society 92(3): 211–265.
Werdelin L. (1991) – Hyaenas: The rise and fall of a carnivore family. Geologiska Föreningen i Stockholm Förhandlingar, 113(1): 95–95.
Werdelin L. (1999) – Studies of fossil hyaenas: affinities of Lycyaenops rhomboidae Kretzoi from Pestlornic, Hungary. Zoological Journal of the Linnean Society 126: 307–317.
Werdelin L., Drăguşin V., Robu M., Petculescu A., Popescu A., Curran S. & Terhune C.E. (2023) – Carnivora from the Early Pleistocene of Grăunceanu (Olteţ River Valley, Dacian Basin, Romania). Rivista Italiana di Paleontologia e Stratigrafia, 129(3): 457–476.
Werdelin L. & Lewis M. (2001) – A revision of the genus Dinofelis (Mammalia, Felidae). Zoological Journal of the Linnean Society, 132: 147–258.
Werdelin L. & Solounias N. (1990) – Studies of fossil hyaenids: the genus Adcrocuta Kretzoi and the interrelationships of some hyaenid taxa. Zoological Journal of the Linnean Society, 98: 363–386.
Werdelin L. & Solounias N. (1991) – The Hyaenidae: taxonomy, systematics and evolution. Fossils and Strata, 30: 1–104.
Werdelin L., Yamaguchi N., Johnson W.E. & O’Brien S.J. (2010) – Phylogeny and evolution of cats (Felidae). In: MacDonald D.W. & Loveridge A.J. (Eds) – Biology and Conservation of Wild Felids: 59–82. Oxford University Press, Oxford.
Willemsen G.F. (1992) – A revision of the Pliocene and Quaternary Lutrinae from Europe. Scripta Geologica, 101: 1–115.
Wolsan M. (1989) – Drapieżne – Carnivora. In: Kowalski K. (Ed.) – Historia i ewolucja lądowej fauny Polski. Folia Quaternaria, 59–60: 177–197.
Zdansky O. (1924) – Jungtertiäre Carnivoren Chinas. Palaeontologia Sinica, 2: 1–149.
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Copyright (c) 2024 NIKOLAOS KARGOPOULOS, ALBERTO VALENCIANO, PANAGIOTIS KAMPOURIDIS, ŞTEFAN VASILE, LAURENŢIU URSACHI, BOGDAN RĂŢOI
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Accepted 2024-06-05
Published 2024-06-28
