THE VERTEBRATE FAUNA OF THE LOWER PLIOCENE CONTINENTAL DEPOSITS IN THE EBRO DELTA (BAIX EBRE BASIN, NE SPAIN)
DOI:
https://doi.org/10.54103/2039-4942/22792Keywords:
Iberian Peninsula, Ruscinian, amphibians, reptiles, mammals, palaeoecologyAbstract
The Pliocene faunas of the northeastern Iberian Peninsula are poorly known due to the scarcity of deposits from this interval. In this context, the site of Sant Nofre-Campredó (Baix Ebre, Catalonia, Spain), comprising two contemporaneous outcrops belonging to geologically correlated sections (Sant Nofre and Campredó-Via Fèrria), makes a significant contribution to the study of Pliocene faunas in this region. The first vertebrate remains were recovered in the late 1970s in clay deposits close to the Sant Nofre Hill. Later, intensive sampling in the 1980s at the Campredó-Via Fèrria outcrop led to the recovery of more than a hundred fossil specimens, including small vertebrates and large mammals. Here, we revise these collections to expand and update the vertebrate list, better constrain the age of the continental deposits, and characterize the environment and climate around the mouth of the Ebro River during the formation of the site. Up to 24 vertebrate species have been identified at Sant Nofre-Campredó, including amphibians, reptiles, and mammals. The presence of the rodents Paraethomys baeticus Piñero & Verzi, 2020, Apocricetus barrierei (Mein & Michaux 1970), and Occitanomys brailloni Michaux, 1969 implies an early Ruscinian age (MN14 unit) for the two outcrops. Biostratigraphic correlations with other well-dated sites in the Iberian Peninsula refine the estimated age of Sant Nofre-Campredó to approximately 4.6 Ma. The ecological affinities of the identified fauna reveal a landscape dominated by woodland habitats in the surroundings of the site during the Early Pliocene, with presence of open humid meadows and water bodies under warm and moist climatic conditions.
References
Adrover R. (1986) - Nuevas faunas de roedores en el Mio-Plioceno continental de la región de Teruel (España). Interés Bioestratigráfico y Paleoecológico. Instituto de Estudios Turolenses, Teruel, 423 pp.
Adrover R., Mein P. & Moissenet, E. (1988) - Contribución al conocimiento de la fauna de roedores del Plioceno de la región de Teruel. Revista del Instituto de Estudios Turolenses, 79: 89-151.
Adrover R., Mein P. & Moissenet E. (1993a) - Los Sciuridae del Turoliense superior, Rusciniense y Villanyense de la región de Teruel. Paleontologia i Evolució, 26-27: 85-106.
Adrover R., Mein P. & Moissenet E. (1993b) - Roedores de la transición Mio-Plioceno de la región de Teruel. Paleontologia i Evolució, 26-27: 47-84.
Aguilar J.P., Michaux J., Bachelet B., Calvet M. & Faillat J.P. (1991) - Les nouvelles faunes de rongeurs proches de la limit Mio-Pliocene en Roussillon. Implications biostratigraphiques et biogéographiques. Palaeovertebrata, 20: 147-174.
Aguirre J., Belaústegui Z., Domènech R., de Gibert J.M. & Martinell, J. (2014) - Snapshot of a lower pliocene dendropoma reef from sant onofre (Baix Ebre Basin, Tarragona, NE Spain). Palaeogeography, Palaeoclimatology, Palaeoecology, 395: 9-20.
Agustí J. & Gibert J. (1985) - Roedores y logomorfos (Mammalia) del Plioceno de San Onofre (Baix Ebre, NE de España). Paleontologia i Evolució, 19: 57-60.
Agustí J., Anadón P. & Julià Brugués R. (1983) - Nuevos datos sobre el Plioceno del Baix Ebre. Aportación a la correlación entre las escalas marina y continental. Acta Geológica Hispánica, 18: 123-130.
Agustí J., Cabrera L., Garcés M., Krijgsman W., Oms O. & Parés J.M. (2001) - A calibrated mammal scale for the Neogene of Western Europe. State of the art. Earth-Science Reviews, 52: 247-260.
Agustí J., Garcés M. & Krijgsman W. (2006) - Evidence for African-Iberian exchanges during the Messinian in the Spanish mammalian record. Palaeogeography, Palaeoclimatology, Palaeoecology, 238: 5-14.
Agustí J., Chochisvili G., Lozano-Fernández I., Furió M., Piñero P. & de Marfà R. (2022) - Small mammals (Insectivora, Rodentia, Lagomorpha) from the Early Pleistocene hominin bearing site of Dmanisi (Georgia). Journal of Human Evolution, 170: 103238.
Aldana Carrasco E.J. (1991) - Roedores Esciuromorfos del Neógeno de Cataluña. Unpublished PhD dissertation, Universitat Autònoma de Barcelona, Bellaterra, 304 pp.
Aldana Carrasco E.J. (1992) - Los Castoridae (Rodentia, Mammalia) del Neógeno de Cataluña (España). Treballs del Museu de Geologia de Barcelona, 2: 99-141.
Alcalá L. (1994) - Macromamíferos neógenos de la fosa de Alfambra-Teruel. Unpublished PhD dissertation, Instituto de Estudios Turolenses-Museo Nacional de Ciencias Naturales, Teruel, 554 pp.
Alcalá L. & Morales J. (1997) - A primitive caprine from the Upper Vallesian of La Roma 2 (Alfambra, Teruel, Aragón, Spain). Comptes Rendus des Séances de l’Académie des Sciences de Paris, 324: 947-953.
Alcalá L. & Morales J. (2006) - Antilopinae (Bovidae, Mammalia) from the Lower Pliocene of Teruel Basin (Spain). Estudios Geológicos, 62: 559-570.
Angelone C. & Rook L. (2012) - Late Neogene and Quaternary lagomorphs from Tuscany: a revision based on specimens in Basel Naturhistorisches Museum and Florence University collections. Swiss Journal of Palaeontology, 131: 127-145.
Angelone C. & Sesé C. (2009) - New characters for species discrimination within the genus Prolagus (Ochotonidae, Lagomorpha, Mammalia). Journal of Paleontology, 83: 80-88.
Arasa A. (1990) - El Terciario del Baix Ebre: aportaciones estratigráficas y sedimentológicas. Acta Geológica Hispánica, 25: 271-288.
Arasa-Tuliesa A. & Cabrera L. (2018) - Neogene-Quaternary onshore record in the lower Ebro river incised palaeovalley (Ebro margin, Catalan Coastal Range, NE Iberia). Geologica Acta, 16: 265-292.
Augé M. (2003) - La faune de Lacertilia (Reptilia, Squamata) de l’Eocène de Prémontré (Bassin de Paris, France). Geodiversitas, 25: 539-574.
Augé M. (2005) - Evolution des lézards du Paléogène en Europe. Mémoires du Muséum national d’Histoire naturelle Paris, 192: 1-369.
Bachelet B. (1990) - Muridae et Arvicolidae (Rodentia, Mammalia) du Pliocène du Sud de la France: systématique, évolution, biochronologie. Unpublished PhD dissertation, Université de Montpellier II, Montpellier, 211 pp.
Bachelet B. & Castillo Ruiz C. (1990) - Radiation évolutive et lignées chez les Stephanomys (Rodentia, Mammalia), muridés dominants du Pliocène d’Europe sud-occidentale. Comptes Rendus des Séances de l’Académie des Sciences de Paris, 311: 493-499.
Bachmayer F. & Wilson R.W. (1978) - A second contribution to the small mammal fauna of Kohfidisch, Austria. Annalen des Naturhistorischen Museums in Wien, 81: 129-161.
Bailon S. (1999) - Différenciation ostéologique des anoures (Amphibia, Anura) de France. In: Desse J. & Desse-Berset N. (Eds) - Fiches d’ostéologie animale pour l’Archéologie, Série C Varia: 3-41. Centre de Recherches Archéologiques, CNRS.
Bailon S. (2010) - Quelonios fósiles del yacimiento de Barranco León (Pleistoceno Inferior, Orce, Granada, España). In: Toro I., Martínez-Navarro B. & Agustí J. (Eds) - Ocupaciones humanas en el pleistoceno inferior y medio de la Cuenca de Guadix-Baza: 185-195. Consejería de Cultura, Junta de Andalucía.
Barahona F. & Barbadillo L.J. (1997) - Identification of some Iberian lacertids using skull characters. Revista Española de Herpetología, 11: 47-62.
Barisone G. (2004) - The fossil record of the genus Castor (Castoridae, Rodentia) in the Neogene and Quaternary of Central and Western Europe, with particular reference to Italy. Unpublished PhD dissertation, Università di Roma Tre, Roma.
Barisone G., Argenti P. & Kotsakis T. (2006) - Plio-Pleistocene evolution of the genus Castor (Rodentia, Mammalia) in Europe: C. fiber plicidens of Pietrafitta (Perugia, Central Italy). Geobios, 39: 757-770.
Bataller J.R. & López M. (1929) - Mapa Geológico Nacional. E. 1:50.000, Hoja nº522, Tortosa, 1ª serie. IGME, 61 pp.
Batbold J., Batsaikhan N., Shar S., Hutterer R., Kryštufek B., Yigit N., Mitsainas G. & Palomo L. (2021) - Castor fiber (amended version of 2016 assessment). The IUCN Red List of Threatened Species 2021, e.T4007A197499749.
Bessais E. & Cravatte J. (1988) - Les écosystèmes végétaux Pliocènes de catalogne méridionale. Variations latitudinales dans le domaine nord-ouest méditerranéen. Geobios, 21: 49-63.
Bessedik M. & Cabrera L. (1985) - Le couple récif-mangrove à Sant Pau d’Ordal (Vallès-Pénédès, Espagne), témoin du maximum transgressif en Méditerranée nord occidentale (Burdigalian supérieur – Langhien inférieur). Newsletter on Stratigraphy, 14: 20-35.
Bibi F. & Güleç E.S. (2008) - Bovidae (Mammalia: Artiodactyla) from the late Miocene of Sivas, Turkey. Journal of Vertebrate Paleontology, 28: 501-519.
Biton R., Geffen E., Vences M., Cohen O., Bailon S., Rabinovich R., Malka Y., Oron T., Boistel R., Brumfeld V. & Gafny S. (2013) - The rediscovered Hula painted frog is a living fossil. Nature Communications, 4: 1959.
Biton R., Boistel R., Rabinovich R., Gafny S., Brumfeld V. & Bailon S. (2016) - Osteological observations on the alytid anura Latonia nigriventer with comments on functional morphology, biogeography, and evolutionary history. Journal of Morphology, 277: 1131-1145.
Blain H.-A., Bailon S., Cuenca-Bescos G., Arsuaga J.L., Bermúdez de Castro J.M. & Carbonell E. (2009) - Long-term climate record inferred from early–middle Pleistocene amphibian and squamate reptile assemblages at the Gran Dolina Cave, Atapuerca, Spain. Journal of Human Evolution, 56: 55-65.
Blain H.-A., Agustí J., López-García J.M., Haddoumi H., Aouraghe H., El Hammouti K., Pérez-González A., Chacon M.G. & Sala R. (2013) - Amphibians and squamate reptiles from the Late Miocene (Vallesian) of Eastern Morocco (Guefaït-1, Jerada Province). Journal of Vertebrate Paleontology, 33: 804-816.
Blain H.-A., Delfino M., Berto C. & Arzarello M. (2016) - First record of Pelobates syriacus (Anura, Amphibia) in the early Pleistocene of Italy. Palaeobiodiversity and Palaeoenvironments, 96: 111-124.
Blain H.-A., Prikryl T., Martínez-Monzón A., Linares-Martín A., Macaluso L., Moreno-Ribas E., Agustí J., Campeny G. & Gómez de Soler B. (2022) - Amphibians and reptiles diversity in the Pliocene Camp dels Ninots maar (Spain). The 6th International Palaeontological Congress “From Gondwana to Laurasia”. Khon Kaen, Thailand, 477 pp.
Blain H.-A., Přikryl T., Piñero P., Sánchez-Bandera C., Martínez-Monzón A. & Fagoaga A. (2024) - Small vertebrates from the Late Pliocene Las Higueruelas locality of central Spain with new biochronological and palaeoecological inferences. Palaeogeography, Palaeoclimatology, Palaeoecology, 635: 111929.
Böhme M. (2010) - Ectothermic vertebrates (Actinopterygii, Allocaudata, Urodela, Anura, Crocodylia, Squamata) from the Miocene of Sandelzhausen (Germany, Bavaria) and their implications for environment reconstruction and palaeoclimate. Paläontologische Zeitschift, 84: 3-41.
Böhme M. & Ilg A. (2003) - fosFARbase [accessed 2024 January]. www.wahrestaerke.com/.
Böhme W., Roček Z. & Špinar Z.V. (1982) - On Pelobates decheni Troschel, 1861, and Zaphrissa eurypelis Cope, 1866 (Amphibia: Salientia: Pelobatidae) from the early Miocene of Rott near Bonn, West Germany. Journal of Vertebrate Paleontology, 2: 1-7.
Bons J. & Geniez P. (1996) - Amphibiens et reptiles du Maroc (Sahara Occidental compris), Atlas biogéographique. Asociación Herpetológica Española, Barcelona, 320 pp.
Buffrénil de V., Dauphin Y., Rage J.-C. & Sire J.-Y. (2011) - An enamel-like tissue, osteodermine, on the osteoderms of a fossil anguid (Glyptosaurinae) lizard. Comptes Rendus Palevol, 10: 427-437.
Cai C. & Qiu Z. (1993) - Murid rodents from the late Pliocene of Yangquan and Yuxian, Hebei. Vertebrata PalAsiatica, 31: 267-293.
Camaiti M., Villa A., Wencker L.C.M., Bauer A.M., Stanley E.L. & Delfino M. (2019) - Descriptive osteology and patterns of limb loss of the European limbless skink Ophiomorus punctatissimus (Squamata, Scincidae). The Anatomical Record, 235: 313-345.
Canerot J., Fonollosa Mr. J. & Wallez M.J. (1975) - Identification du Miocène inférieur-moyen dans la basse vallée de l’Ebre. Conséquences sur l’age de la tectorogènese alpine aux confins des Chaines Ibérique et Catalane (Espagne). Comptes Rendus des Séances de l’Académie des Sciences de Paris, 280: 1951-1954.
Carrión J.S., Barrón E., Postigo Mijarra J.M., Casas-Gallego M., Munuera M., Ochando J., Jiménez-Moreno G., Amorós G., Altolaguirre Y., Vieira M., Tosal A., Moreno-Domínguez R., Rivas-Carballo R., Valle-Hernández M., Alonso-Gavilán G., Fernández S., Verdú M., Arroyo J., Di Rita F., Magri D., Gómez-Rodríguez M., Sánchez-Giner V., Amorós A., Fernández-Díaz M., Reyes P.P. (2022) - Paleoflora y paleoevegetación ibérica I: Paleoceno-Plioceno, Paleoflora y paleovegetación ibérica. Ministerio de Ciencia e Innovación y Fundación Séneca, Murcia, 225 pp.
Casanovas-Vilar I. & Agustí J. (2007) - Ecogeographical stability and climate forcing in the Late Miocene (Vallesian) rodent record of Spain. Palaeogeography, Palaeoclimatology, Palaeoecology, 248: 169-189.
Casanovas-Vilar I. & Alba D.M. (2011) - The never-ending problem of Miocene beaver taxonomy. Acta Palaeontologica Polonica, 56: 217-220.
Casanovas-Vilar I., Alba D.M., Almécija S., Robles J.M., Galindo J. & Moyà-Solà S. (2008) - Taxonomy and paleobiology of the genus Chalicomys Kaup, 1832 (Rodentia, Castoridae), with the description of a new species from Abocador de Can Mata (Vallès-Penedès Basin, Catalonia, Spain). Journal of Vertebrate Paleontology, 28: 851-862.
Casanovas-Vilar I., van Dam J.A., Moyà-Solà S. & Rook L. (2011) - Late Miocene insular mice from the Tusco-Sardinian palaeobioprovince provide new insights on the palaeoecology of the Oreopithecus faunas. Journal of Human Evolution, 61: 42-49.
Casanovas-Vilar I., Almécija S. & Alba D.M. (2015) - Late Miocene flying squirrels from Can Llobateres 1 (Vallès-Penedès Basin, Catalonia): systematics and palaeobiogeography. Palaeobiodiversity and Palaeoenvironments, 95: 353-372.
Castillo C. (1990) - Paleocomunidades de micromamíferos de los yacimientos kársticos del Neógeno Superior de Andalucía Oriental. Unpublished PhD Dissertation, Universidad de Granada, Granada, 255 pp.
Castillo Ruiz C. (1992) - Paraethomys belmezensis nov. sp. (Rodentia, Mammalia) du Pliocène de Córdoba (Espagne). Geobios, 25: 775-780.
Čerňanský A. & Klembara J. (2017) - A skeleton of Ophisaurus (Squamata: Anguidae) from the middle Miocene of Germany, with a revision of the partly articulated postcranial material from Slovakia using micro-computed tomography. Journal of Vertebrate Paleontology, 37: e1333515.
Čerňanský A., Yaryhin O., Ciceková J., Werneburg I., Hain M. & Klembara J. (2019) - Vertebral comparative anatomy and morphological differences in anguine lizards with a special reference to Pseudopus apodus: Vertebral column and ribs of Pseudopus apodus. The Anatomical Record, 302: 232-257.
Cherin M., Alba D.M., Crotti M., Menconero S., Moullé P.-E., Sorbelli L. & Madurell-Malapeira J. (2020) - The post-Jaramillo persistence of Sus strozzii (Suidae, Mammalia) in Europe: New evidence from the Vallparadís Section (NE Iberian Peninsula) and other coeval sites. Quaternary Science Reviews, 2,
: 106234.
Colombero S., Pavia G. & Carnevale G. (2014) - Messinian rodents from Moncucco Torinese, NW Italy: palaeobiodiversity and biochronology. Geodiversitas, 36: 421-475.
Cordy J.M. (1976) - Essai sur la microévolution du genre Stephanomys (Rodentia, Muridae). Unpublished PhD dissertation. University of Liège, 351 pp.
Cox P.G. & Baverstock H. (2016) - Masticatory muscle anatomy and feeding efficiency of the American Beaver, Castor canadensis (Rodentia, Castoridae). Journal of Mammal Evolution, 23: 191-200.
Crusafont Pairó M., De Villalta J.F. & Bataller J.R. (1948) - Los castores fósiles de España. Boletín del Instituto Geológico y Minero de España, 61: 321-449.
Cuenca-Bescós G. (1988) - Revisión de los Sciuridae del Aragoniense y del Rambliense en la fosa de Calatayud-Montalbán. Scripta Geologica, 87: 1-116.
Cuenca-Bescós G., Rosell Ardévol J., Morcillo-Amo Á., Galindo-Pellicena M.Á., Santos E. & Moya-Costa R. (2017) - Beavers (Castoridae, Rodentia, Mammalia) from the Quaternary sites of the Sierra de Atapuerca, in Burgos, Spain. Quaternary International, 433: 263-277.
Daams R. (1981) - The dental pattern of the Dormice Dryomys, Myomimus, Microdyromys and Peridyromys. Utrecht Micropaleontological Bulletin, Special Publication, 3: 1-115.
De Bruijn H. (1974) - The Ruscinian rodent succession in Southern Spain and its implications for the biostratigraphic correlation of Europe and North Africa. Senckenbergiana Lethaea, 55: 435-443.
de Gibert J.M. & Martinell J. (1996) - Trace fossil assemblages and their palaeoenvironmental significance in the Pliocene marine deposits of the Baix Ebre (Catalonia, NE of Spain). Géologie Méditerranéenne, 23: 211-225.
de Gibert J.M. & Martinell J. (1998) - Ichnofabrics of the Pliocene marginal marine basins of the northwestern Mediterranean. Revista de la Sociedad Geológica de España, 11: 43-56.
de Gibert J.M., Martinell J. & Domènech R. (1998) - Entobia ichnofacies in fossil rocky shores, lower Pliocene, northwestern Mediterranean. Palaios, 13: 476-487.
de Giuli C. (1989) - The rodents of the Brisighella latest Miocene fauna. Bollettino Della Societa Paleontologica Italiana, 28: 197-212.
Escoriza D. & Comas M. (2015) - Is Hyalosaurus koellikeri a true forest lizard? Herpetological Conservation and Biology, 10: 610-620.
Estes R. (1981) - Gymnophiona, Caudata. Handbuch der Paläoherpetologie, 2: 1-115.
Fauquette S., Suc J.-P., Guiot J., Diniz F., Feddi N., Zheng Z., Bessais E. & Drivaliari A. (1999) - Climate and biomes in the West Mediterranean area during the Pliocene. Palaeogeography, Palaeoclimatology, Palaeoecology, 152: 15-36.
Fejfar O. & Storch, G. (1990) - Eine pliozäne (ober-ruscinische) Kleinsäugerfauna aus Gundersheim, Rheinhessen. I. Nagetiere: mammalia, Rodentia. Senckenbergiana Lethaea, 71: 139-184.
Fleta J., Arasa A. & Escuer J. (1991) - El Neógeno del Empordà y Baix Ebre (Catalunya): estudio comparativo. Acta Geológica Hispánica, 26: 159-171.
Font y Sagué N. (1905). Nota sobre la preséncia del terreno pliocénich en la comarca de Tortosa. Butlletí de la Institució Catalana d'Història Natural, 2: 59-61.
Freudenthal M. (2004) - Gliridae (Rodentia, Mammalia) from the Eocene and Oligocene of the Sierra Palomera (Teruel, Spain). Treballs del Museu de Geologia de Barcelona, 12: 97-173.
Freudenthal M., Mein P. & Martín-Suárez E. (1998) - Revision of Late Miocene and Pliocene Cricetinae (Rodentia, Mammalia) from Spain and France. Treballs del Museu de Geologia de Barcelona, 7: 11-93.
Friant M. (1964) - An European Castor of the Quaternary era, Castor plicidens F. Maj. Journal of the Palaeontological Society of India, 4: 75-77.
Furió M. & Agustí J. (2017) - Latest Miocene insectivores from Eastern Spain: Evidence for enhanced latitudinal differences during the Messinian. Geobios, 50: 123-140.
Furió M., Casanovas-Vilar I. & van den Hoek Ostende L.W. (2011) - Predictable structure of Miocene insectivore (Lipotyphla) faunas in Western Europe along a latitudinal gradient. Palaeogeography, Palaeoclimatology, Palaeoecology, 304: 219-229.
Furió M., Gibert L., Ferràndez C. & Sevilla P. (2015) - The insectivores (Soricidae, Erinaceidae; Eulipotyphla; Mammalia) from Cueva Victoria (Early Pleistocene, Murcia, Spain). Neues Jahrbuch für Geologie und Paläontologie, 275: 151-161.
Furió M., van den Hoek Ostende L.W., Agustí J. & Minwer-Barakat R. (2018) - Evolution of the insectivore assemblages (Eulipotyphla, Mammalia) in Spain and their relation with Neogene and Quaternary climatic changes. Ecosistemas, 27: 38-51.
Garcés M., Krijgsman W. & Agustí J. (2001) - Chronostratigraphic framework and evolution of the Fortuna basin (Eastern Betics) since the Late Miocene. Basin Research, 13: 199-217.
García-Alix A., Minwer-Barakat R., Martín-Suárez E. & Freudenthal M. (2007) - New data on Mio-Pliocene Sciuridae (Rodentia, Mammalia) from southern Spain. Comptes Rendus Palevol, 6: 269-279.
García-Alix A., Minwer-Barakat R., Martín-Suárez E. & Freudenthal M. (2008a) - Muscardinus meridionalis sp. nov., a new species of Gliridae (Rodentia, Mammalia) and its implications for the phylogeny of Muscardinus. Journal of Vertebrate Paleontology, 28: 568-573.
García-Alix A., Minwer-Barakat R., Martín-Suárez E. & Freudenthal M. (2008b) - Cricetidae and Gliridae (Rodentia, Mammalia) from the Miocene and Pliocene of southern Spain. Scripta Geologica 136, 1-37.
García-Alix A., Minwer-Barakat R., Martín-Suárez E. & Freudenthal M. (2008c) - Muridae (Rodentia, Mammalia) from the Mio-Pliocene boundary in the Granada Basin (southern Spain). Biostratigraphic and phylogenetic implications. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, 248: 183–215.
García-Alix A., Minwer-Barakat R., Martín-Suárez E. & Freudenthal M. (2008d) - Biostratigraphy and sedimentary evolution of Late Miocene and Pliocene continental deposits of the Granada Basin (southern Spain). Lethaia, 41: 431-446.
García-Alix A., Minwer-Barakat R., Martín-Suárez E., Freudenthal M. & Martín J.M. (2008e) - Late Miocene–Early Pliocene climatic evolution of the Granada Basin (southern Spain) deduced from the paleoecology of the micromammal associations. Palaeogeography, Palaeoclimatology, Palaeoecology, 265: 214-225.
Garcia-Ibaibarriaga N., Arrizabalaga Á., Iriarte-Chiapusso M.J., Rofes J. & Murelaga X. (2015) - The return to the Iberian Peninsula: first Quaternary record of Muscardinus and a palaeogeographical overview of the genus in Europe. Quaternary Science Reviews, 119: 106-115.
Gibert J., Agustí J. & Moyà S. (1980) - Nuevos datos sobre la bioestratigrafía del Ampurdán. Boletín Geológico y Minero, 91: 705-712.
Gignoux M. & Fallot P. (1922) - Le pliocène marin sur les côtes méditerranéeenes d'Espagne. Comptes Rendus des Séances de l’Académie des Sciences de Paris, 249: 2351-2353.
Gmelig-Meyling C. & Michaux J. (1973) - Le genre Stephanomys Schaub, 1938; son évolution au Pliocène supérieur. Comptes Rendus des Séances de l’Académie des Sciences de Paris, 277: 1441-1442.
Gomes Rodrigues H., Renaud S., Charles C., Le Poul Y., Solé F., Aguilar J.-P., Michaux J., Tafforeau P., Headon D., Jernvall J. & Viriot L. (2013) - Roles of dental development and adaptation in rodent evolution. Nature Communications, 4: 2504.
Gómez-Cano A.R., Hernandez-Fernandez M. & Álvarez-Sierra M.Á. (2013) - Dietary ecology of Murinae (Muridae, Rodentia): a geometric morphometric approach. PLoS One, 8: e79080.
Gómez-Cano A.R., Kimura Y., Blanco F., Menéndez I., Álvarez-Sierra M.A. & Hernández-Fernández M. (2017) - Ecomorphological characterization of murines and non-arvicoline cricetids (Rodentia) from south-western Europe since the latest Middle Miocene to the Mio-Pliocene boundary (MN 7/8–MN13). PeerJ, 5: e3646.
Gómez de Soler B., Campeny Vall-llosera G., van der Made J., Oms O., Agustí J., Sala R., Blain H.-A., Burjachs F., Claude J., García Catalan S., Riba D. & Rosillo R. (2012) - A new key locality for the Pliocene vertebrate record of Europe: the Camp dels Ninots maar (NE Spain). Geologica Acta, 10: 1-17.
Guimerà J. & Álvaro M. (1990) - Structure et évolution de la compression alpine dans la Chaine ibérique et la Chaîne côtière catalane (Espagne). Bulletin de la Société Géologique de France, 6: 339-340.
Haywood A.M., Dowsett H.J., Robinson M.M., Stoll D.K., Dolan A.M., Lunt D.J., Otto-Bliesner B. & Chandler M.A. (2011) - Pliocene Model Intercomparison Project (PlioMIP): experimental design and boundary conditions (Experiment 2). Geoscientific Model Development, 4: 571-577.
Heller F. (1936) - Eine oberpliozäne Wirbeltierfauna aus Rheinhessen. Neues Jahrbuch für Mineralogie, Geologie und Palaöntologie, Abteilung B, 76: 99-160.
Hellmund M. & Böhme W. (1987) - Zweiter Fund eines vollständiges Exemplares von Chelotriton paradoxus Pomel, 1853 aus dem Oberoligozän von Rott bei Bonn. Salamandra, 23: 142-152.
Hernández-Fernández M. & Peláez-Campomanes P. (2003) - Ecomorphological characterization of Murinae and hypsodont Cricetidae (Rodentia) from the Iberian Plio-Pleistocene. Coloquios de Paleontología Vol Ext., 1: 237-251.
Hervet S. (2000) - Tortues du Quaternaire de France: critères de détermination, répartitions chronologique et geographique. Mésogée, 58: 3-47.
Hoffstetter R. (1962) - Observations sur les ostéodermes et la classification des Anguidés actuels et fossiles (Reptiles, Sauriens). Bulletin du Muséum National d’Histoire Naturelle, 34: 149-157.
Hoffstetter P. & Gasc J.P. (1969) - Vertebrae and ribs of modern reptiles. In: Gans C. (Ed) - Biology of the Reptilia. Vol. 1: Morphology A: 201-210. Academic Press, New York.
Hordijk K. & De Bruijn H. (2009) - The succession of rodent faunas from the Mio/Pliocene lacustrine deposits of the Florina-Ptolemais-Servia Basin (Greece). Hellenic Journal of Geosciences, 44: 21-103.
Hossini S. (2000) - Un nouveau discoglossidé (Amphibia, Anura) dans le Miocène supérieur du gisement marocain “Oued Zra”. Comptes Rendus des Séances de l’Académie des Sciences de Paris, 331: 621-626.
Hugueney M. (2004) - Les grands rongeurs du Pliocène supérieur de Saint-Vallier (Drôme, France): Castoridae, Hystricidae (Mammalia, Rodentia). Geobios, 37: 126-132.
Hugueney M. & Mein P. (1965) - Lagomorphes et rongeurs du Néogène de Lissieu (Rhône). Travaux et Documents des Laboratoires de Géologie de Lyon, 12: 109-124.
Iannucci A., Cherin M., Sorbelli L. & Sardella R. (2021) - Suidae transition at the Miocene-Pliocene boundary: a reassessment of the taxonomy and chronology of Propotamochoerus provincialis. Journal of Mammalian Evolution, 28: 323-335.
Ivanov M. (2002) - The oldest known Miocene snake fauna from Central Europe: Merkur-North locality, Czech Republic. Acta Palaeontologica Polonica, 47: 513-534.
Ivanov M. (2008) - Early Miocene Amphibians (Caudata, Salientia) from the Mokrá-Western Quarry (Czech Republic) with comments on the evolution of Early Miocene amphibian assemblages in Central Europe. Geobios, 41: 465-492.
Jiménez-Moreno G., Fauquette S. & Suc J.-P. (2010) - Miocene to Pliocene vegetation reconstruction and climate estimates in the Iberian Peninsula from pollen data. Review of Palaeobotany and Palynology, 162: 403-415.
Knitlová M. & Horáček I. (2017) - Late Pleistocene-Holocene paleobiogeography of the genus Apodemus in central Europe. PLoS One, 12: e0173668.
Köhler M., Moyà-Solà S. & Morales J. (1995) - The vertebrate locality Maramena (Macedonia, Greece) at the Turolian–Ruscinian boundary (Neogene). 15. Bovidae and Giraffidae (Artiodactyla, Mammalia). Münchner Geowissenschaftliche Abhandlungen A, 25: 167-180.
Lizana M. (1997) - Pelobates cultripes. In: Pleguezuelos J.M. & Martínez-Rica J.P. (Eds) - Distribución y Biogeografia de los anfibios y reptiles de España. Monografias Revista Española de Herpetología 3: 140-142. Universidad de Granada, Granada.
Llenas M. & Lorente À.G. (2002) - Els petits mamífers del Neògen i del Quaternari inferior. Quaderns, 23: 29-42.
López-García J.M., Piñero P., Agustí J., Furió M., Galán J., Moncunill-Solé B., Ruiz-Sánchez F.J., Blain H.-A., Sanz M. & Daura J. (2024) - Chronological context, species occurrence, and environmental remarks on the Gelasian site Pedrera del Corral d’en Bruach (Barcelona, Spain) based on the small-mammal associations. Historical Biology, 36: 657-676.
Magné J. (1978) - Études microstratigraphiques sur le Néogène de la Méditerranée nord-occidentale: Les bassins néogènes catalans. Éditions du CNRS, 259 pp.
Magné J. & Viallard P. (1977) - L'evolution tectonique alpine dans la Chaine Ibérique: le Tertiaire marin aus abords du delta de l'Ebre et du Pliocène post-tectorogénique. Comptes Rendus des Séances de l’Académie des Sciences de Paris, 285: 1275-1278.
Maldonado A. (1972) - El Delta del Ebro. Estudio sedimentológico y estratigráfico. Boletín de Estratigrafía, 1: 1-486.
Maldonado-Garrido E., Piñero P. & Agustí J. (2017) - A catalogue of the vertebrate fossil record from the Guadix-Baza Basin (SE Spain). Spanish Journal of Palaeontology, 32: 207-235.
Mansino S., Fierro I., Ruiz-Sánchez F.J. & Montoya P. (2013) - The fossil rodent faunas of the localities Alcoy 2C and 2D (Alcoy Basin, Spain). Implications for dating the classical locality of Alcoy-Mina. Journal of Iberian Geology, 39: 261-284.
Mansino S., Ruiz-Sánchez F.J., Freudenthal M. & Montoya P. (2014) - A new approach to the Late Miocene–Early Pliocene forms of the genus Apocricetus. Apocricetus alberti (Rodentia, Mammalia) from Venta del Moro (Cabriel Basin, Spain). Proceedings of the Geologists’ Association, 125: 392-405.
Mansino S., Ruiz-Sánchez F.J., de Luque L., Montoya P., Gibert L., Morales J., Abellá J., Crespo V.D. & Scott J.R. (2015a) - First Early Pliocene micromammal faunas from the Venta del Moro area (Cabriel Basin, Spain): new data on the Messinian dispersal of Debruijnimys. Journal of Iberian Geology, 41: 273-293.
Mansino S., Fierro I., Montoya P. & Ruiz-Sánchez F.J. (2015b) - Micromammal faunas from the Mio-Pliocene boundary in the Alcoy Basin (SE Spain): biostratigraphical and palaeoecological inferences. Bulletin of Geosciences, 90: 555-576.
Mansino S., Ruiz-Sánchez F.J., Fierro I. & Montoya P. (2016) - Mio-Pliocene rodent assemblages from Alcoi Forn (Alcoy Basin, Eastern Spain). Biostratigraphical and palaeoclimatical inferences. Historical Biology, 28: 1050-1065.
Mansino S., Crespo V., Montoya P. & Ruiz-Sánchez F.J. (2017) - Muridae from the late Miocene site of Venta del Moro (Eastern Spain). Historical Biology, 29: 677-691.
Mansino S., Ruiz-Sánchez F.J. & Montoya P. (2018) - Sciuridae (Rodentia, Mammalia) from the Late Miocene of Venta del Moro (Cabriel Basin, Spain): biostratigraphical, phylogenetic and palaeoecological inferences. Historical Biology, 30: 297-304.
Martín-Suárez E. (1988) - Sucesiones de micromamíferos en la Depresión de Guadix-Baza (Granada, España). Unpublished PhD dissertation, Universidad de Granada, Granada, 241 pp.
Martín-Suárez E. & Freudenthal M. (1993) - Muridae (Rodentia) from the lower Turolian of Crevillente (Alicante, Spain). Scripta Geologica, 103: 65-118.
Martín-Suárez E. & Mein P. (1991) - Revision of the genus Castillomys (Muridae, Rodentia). Scripta Geologica, 96: 47-81.
Martín-Suárez E. & Mein P. (1998) - Revision of the genera Parapodemus, Apodemus, Rhagamys and Rhagapodemus (Rodentia, Mammalia). Geobios, 31: 87-97.
Martín-Suárez E. & Mein P. (2004) - The late Pliocene locality of Saint-Vallier (Drôme, France). Eleven Micromammals. Geobios, 37: 115-125.
Martín-Suárez E., Oms O., Freudenthal M., Agustí J. & Parés J.M. (1998) - Continental Mio-Pliocene transition in the Granada Basin. Lethaia, 31: 161-166.
Martín-Suárez E., Freudenthal M. & Civis J. (2001) - Rodent paleoecology of the Continental Upper Miocene of Crevillente (Alicante, SE Spain). Paleogeography, Paleoclimatology, Paleoecology, 165: 349-356.
Martinell J. (1988) - An overview of the marine Pliocene of NE Spain. Géologie Méditerranéenne, 15: 227-233.
Martinell J. & Domènech R. (1984) - Malacofauna del Plioceno de Sant Onofre (Baix Ebre; Tarragona). Iberus, 4: 1-17.
Martinell J. & Domènech R. (1995) - Bioerosive structures on the Pliocene rocky shores of Catalonia (Spain). Revista Española de Paleontología, 10: 37-44.
Maul L., Masini F., Abbazzi L. & Turner A. (1998) - The use of different morphometric data for absolute age calibration of some South-and Middle European arvicolid populations. Palaeontographia Italica, 85: 111-151.
Mein P. & Michaux J. (1970) - Un nouveau stade dans l’évolution des rongeurs pliocènes de l’Europe sud-occidentale. Comptes Rendus des Séances de l’Académie des Sciences de Paris, 270: 2780-2783.
Mein P. & Freudenthal M. (1971) - Une nouvelle classification des Cricetidae (Mammalia, Rodentia) du Tertiaire de l’Europe. Scripta Geologica, 2: 1-37.
Mein P., Moissenet E. & Adrover R. (1990) - Biostratigraphie du Néogène Supérieur du bassin de Teruel. Paleontologia i Evolució, 23: 121-139.
Michaux J. (1969) - Muridae (Rodentia) du Pliocène supérieur d’Espagne et du midi de la France. Paleovertebrata, 3: 1-25.
Michaux J. (1967) - Origine du dessin dentaire Apodemus (Rodentia, Mammalia). Comptes Rendus des Séances de l’Académie des Sciences de Paris, 265: 711-714.
Michaux J. (1971) - Muridae (Rodentia) néogènes d’Europe sud-occidentale. Evolution et rapports avec les formes actuelles. Paléobiologie Continentale, 2: 1-67.
Miller K.G., Browning J.V., Schmelz W.J., Kopp R.E., Mountain G.S. & Wright J.D. (2020) - Cenozoic sea-level and cryospheric evolution from deep-sea geochemical and continental margin records. Science Advances, 6: eaaz1346.
Minwer-Barakat R. (2005) - Roedores e insectívoros del Turoliense Superior y el Plioceno del sector central de la cuenca de Guadix. Unpublished PhD dissertation, Universidad de Granada, 535 pp.
Minwer-Barakat R., García-Alix A., Martín-Suárez E. & Freudenthal M. (2005) - Muridae (Rodentia) from the Pliocene of Tollo de Chiclana (Granada, South-Eastern Spain). Journal of Vertebrate Paleontology, 25: 426-441.
Minwer-Barakat R., García-Alix A., Agustí J., Martín-Suárez E. & Freudenthal M. (2009a) - The micromammal fauna from Negratín-1 (Guadix Basin, southern Spain): new evidence of African-Iberian mammal exchanges during the Late Miocene. Journal of Paleontology, 83: 854-879.
Minwer-Barakat R., García-Alix A., Martín-Suárez E. & Freudenthal M. (2009b) - Late Turolian micromammals from Rambla de Chimeneas-3: considerations on the oldest continental faunas from the Guadix Basin (southern Spain). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, 251: 95-108.
Minwer-Barakat R., García-Alix A. & Martín-Suárez E. (2011) - Validation of the species Stephanomys progressus, a murid (Rodentia) from the Early Pleistocene of Spain. Journal of Paleontology, 85: 392-394.
Miralles A., Hipsley C.A., Erens J., Gehara M., Rakotoarison A., Glaw F., Müller J. & Vences M. (2015) - Distinct patterns of desynchronized limb regression in Malagasy scincine lizards (Squamata, Scincidae). PLoS One, 10: e0126074.
Montenat C. & De Bruijn H. (1976) - The Ruscinian rodent faunule from La Juliana (Murcia); its implication for correlation of continental and marine biozones. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen B, 79: 245-255.
Montuire S., Maridet O. & Legendre S. (2006) - Late Miocene–Early Pliocene temperature estimates in Europe using rodents. Palaeogeography, Palaeoclimatology, Palaeoecology, 238: 247-262.
Montoya P., Ginsburg L., Alberdi M.T., Van der Made J., Morales J. & Soria M.D. (2006) - Fossil large mammals from the early Pliocene locality of Alcoy (Spain) and their importance in biostratigraphy. Geodiversitas, 28: 137-173.
Morales J. (1984) - Venta del Moro: Su macrofauna de mamíferos, y biostratigrafía continental del Mioceno terminal mediterráneo. Unpublished PhD Dissertation, Universidad Complutense de Madrid.
Ogg J.G. (2020) - Chapter 5 - Geomagnetic Polarity Time Scale. In: Gradstein F.M., Ogg J.G., Schmitz M.D. & Ogg G.M. (Eds) - Geologic Time Scale 2020: 159-192. Elsevier, Amsterdam.
Oms O., Dinarès-Turell J., Agustı́ J. & Parés J.M. (1999) - Refinements of the European mammal biochronology from the magnetic polarity record of the Plio-Pleistocene Zújar section, Guadix-Baza basin, SE Spain. Quaternary Research, 51: 94-103.
Opdyke N., Mein P., Lindsay E., Pérez-Gonzales A., Moissenet E. & Norton V.L. (1997) - Continental deposits, magnetostratigraphy and vertebrate paleontology, late Neogene of Eastern Spain. Palaeogeography, Palaeoclimatology, Palaeoecology, 133: 129-148.
Pagani M., Liu Z., LaRiviere J. & Ravelo A.C. (2010) - High Earth-system climate sensitivity determined from Pliocene carbon dioxide concentrations. Nature Geoscience, 3: 27-30.
Pasquier L. (1974) - Dynamique évolutive d’un sous-genre de Muridae Apodemus (Sylvaemus). Etude biométrique des caractères dentaires de populations fossiles et actuelles d’Europe occidentale. Unpublished PhD dissertation, University Montpellier, France, 176 pp.
Pérez-García A. (2017) - The Iberian fossil record of turtles: An update. Journal of Iberian Geology, 43: 155-191.
Perl R.G.B., Gafny S., Malka Y., Renan S., Woodhams D.C., Rollins-Smith L., Pask J.D., Bletz M.C., Geffen E. & Vences M. (2017) - Natural history and conservation of the rediscovered Hula painted frog, Latonia nigriventer. Contributions to Zoology, 86: 11-37.
Pickford M. (2013) - Re-assessment of the suids from the Sables marins de Montpellier and selection of a lectotype for Sus provincialis Blainville, 1847. Geodiversitas, 35: 655-689.
Pickford M. & Obada T. (2016) - Pliocene suids from Musaitu and Dermenji, Moldova: implications for understanding the origin of African Kolpochoerus Van Hoepen and Van Hoepen, 1932. Geodiversitas, 38: 99-134.
Piñero P. & Agustí J. (2019) - The rodent succession in the Sifón de Librilla section (Fortuna Basin, SE Spain): implications for the Mio-Pliocene boundary in the Mediterranean terrestrial record. Historical Biology, 31: 279-321.
Piñero P. & Agustí J. (2020) - Rodents from Botardo-D and the Miocene–Pliocene transition in the Guadix-Baza Basin (Granada, Spain). Palaeobiodiversity and Palaeoenvironments, 100: 903-920.
Piñero P. & Verzi D.H. (2020) - A new early Pliocene murine rodent from the Iberian Peninsula and its biostratigraphic implications. Acta Palaeontologica Polonica, 65: 719-731.
Piñero P., Agustí J., Blain H.-A. & Laplana C. (2016) - Paleoenvironmental reconstruction of the Early Pleistocene site of Quibas (SE Spain) using a rodent assemblage. Comptes Rendus Palevol, 15: 659-668.
Piñero P., Agustí J., Oms O., Blain H.-A., Laplana C., Ros-Montoya S. & Martínez-Navarro B. (2017a) - Rodents from Baza-1 (Guadix-Baza Basin, southeast Spain): filling the gap of the early Pliocene succession in the Betics. Journal of Vertebrate Paleontology, 37: e1338294.
Piñero P., Agustí J., Oms O., Fierro I., Montoya P., Mansino S., Ruiz-Sánchez F.R., Alba D.M., Alberdi M.T., Blain H.-A., Laplana C., Van der Made J., Mazo A.V., Morales J., Murelaga X., Pérez-García A., Pérez-Valera F., Pérez-Valera J.A., Sevilla P., Soria J.M. & Romero G. (2017b) - Early Pliocene continental vertebrate fauna at Puerto de la Cadena (SE Spain) and its bearing on the marine-continental correlation of the late Neogene of Eastern Betics. Palaeogeography, Palaeoclimatology, Palaeoecology, 479: 102-114.
Piñero P., Agustí J., Furió M. & Laplana C. (2018a) - Rodents and insectivores from the late Miocene of Romerales (Fortuna Basin, Southern Spain). Historical Biology, 30: 336-359.
Piñero P., Agustí J. & Oms O. (2018b) - The late Neogene rodent succession of the Guadix-Baza Basin (south-eastern Spain) and its magnetostratigraphic correlation. Palaeontology, 61: 253-272.
Piñero P., Agustí J., Laborda C., Duval M., Zhao J.-x., Blain H.-A., Furió M., Laplana C., Rosas A. & Sevilla P. (2022) - Quibas-Sima: a unique 1 Ma-old vertebrate succession in southern Iberian Peninsula. Quaternary Science Reviews, 283: 107469.
Piñero P., Martín-Perea D. M., Sevilla P., Agustí J., Blain H.-A., Furió M. & Laplana C. (2023a) - La Piquera in central Iberian Peninsula: A new key vertebrate locality for the Early Pliocene of western Europe. Acta Palaeontologica Polonica, 68: 23-46.
Piñero P., Agustí J. & Ruiz-Sánchez F.J. (2023b) - Influence of climatic variations on the evolution of the European Paraethomys lineage (Rodentia, Muridae). In: Ros-Franch et al. (Eds) - Libro de Resúmenes de las XXXVIII Jornadas SEP. Palaeontological publications 4: 190.
Piñero P., Agustí J., Blain H.-A., Alberdi M.T., Blanco-Lapaz A. & Furió M. (2024a) - A revisit to the Early Pliocene site of Abla (Guadix-Baza Basin, southern Spain). Geobios, 85: 79-89.
Piñero P., López-García J.M., Blain H.-A., Carnevale C., Furió M., Giuntelli P., Luzi E., Macaluso L., Marramàc G., Pal S., Pavia G., Pavia M., Pezzetti C., Rocca M., Sánchez-Bandera C., Villa A. & Delfino M. (2024b) - Multiproxy approach to reconstruct the climate and environment of a new late Middle Pleistocene vertebrate site in northwestern Italy. Palaeogeography, Palaeoclimatology, Palaeoecology, 634: 111935.
Rage J.C. (1988) - The oldest known colubrid snakes. The state of the art. Acta Zoologica Cracoviensia, 31: 457-474.
Renaud S. & Van Dam J. (2002) - Influence of biotic and abiotic environment on dental size and shape evolution in a Late Miocene lineage of murine rodents (Teruel Basin, Spain). Palaeogeography, Palaeoclimatology, Palaeoecology, 184: 163-175.
Renaud S., Benammi M., Jaeger J.-J. (1999) - Morphological evolution of the murine rodent Paraethomys in response to climatic variations (Mio-Pleistocene of North Africa). Paleobiology, 25: 369-382.
Reumer J.W.F. (1984) - Ruscinian and early Pleistocene Soricidae (Insectivora, Mammalia) from Tegelen (The Netherlands) and Hungary. Scripta Geologica, 73: 1-173.
Reumer J.W.F. (1998) - A classification of the fossil and recent shrews. In: Wójcik J.M. & Wolsan M. (Eds) - Evolution of shrews: 5-22. Mammal Research Institute Polish Academy of Sciences, Białowieża.
Rietschel S. & Storch G. (1974) - Aussergewöhnlich erhaltene Waldmäuse (Apodemus atavus Heller, 1936) aus dem Ober-Pliozän von Willershausen am Harz. Senckenbergiana Lethaea, 54: 491-519.
Roček Z. (1994) - Taxonomy and distribution of Tertiary discoglossids (Anura) of the genus Latonia v. Meyer, 1843. Geobios, 27: 717-751.
Roček Z. (2013) - Mesozoic and tertiary anura of Laurasia. Palaeobiodiversity and Palaeoenvironments, 93: 397-439.
Roček Z. & Wuttke M. (2010) - Amphibia of Enspel (late Oligocene, Germany). Palaeobiodiversity and Palaeoenvironments, 90: 321-340.
Ruiz Bustos A., Sesé C., Dabrio C.J., Peña Ruano J.A. & Padial J.M. (1984) - Geología y fauna de micromamíferos del nuevo yacimiento del Plioceno inferior de Gorafe-A (Depresión de Guadix-Baza, Granada). Estudios Geológicos, 40: 231-241.
Rull V., Alba D. & Casanovas-Vilar I. (2024) - Middle Miocene vegetation of the Vallès-Penedès Basin (NE Iberian Peninsula), as inferred from fossil pollen records: State of the art and future prospects. Review of Palaeobotany and Palynology, 321: 105042.
Saarinen J., Oksanen O., Žliobaitė I., Fortelius M., DeMiguel D., Azanza B., Bocherens H., Luzón C., Solano-García J., Yravedra J., Courtenay L.A., Blain H.-A., Sánchez-Bandera C., Serrano-Ramos A., Rodríguez-Alba J.J., Viranta S., Barksy D., Tallavaara M., Oms O., Agustí J. & Jiménez-Arenas J.M. (2021) - Pliocene to Middle Pleistocene climate history in the Guadix-Baza Basin, and the environmental conditions of early Homo dispersal in Europe. Quaternary Science Reviews, 268: 107132.
Sánchez-Bandera C., Oms O., Blain H.-A., Lozano-Fernández I., Bisbal-Chinesta J.F., Agustí J., Saarinen J., Fortelius M., Titton S., Serrano-Ramos A., Luzón C., Solano-García J., Barsky D. & Jiménez-Arenas J.M. (2020) - New stratigraphically constrained palaeoenvironmental reconstructions for the first human settlement in Western Europe: The Early Pleistocene herpetofaunal assemblages from Barranco León and Fuente Nueva 3 (Granada, SE Spain). Quaternary Science Reviews, 243: 106466.
Sanchiz B. (1998) - Encyclopedia of paleoherpetology, part 4: Salientia. Munich, Verlag Dr. Friedrich Pfeil, 275 pp.
Sanz E., Comas P. & Maroto J. (1987) - Inventari paleontològic dels fons del Museu Arqueològic Comarcal de Banyoles procedents de la zona Banyoles-Mata i adjacents. Quaderns del Centre d’Estudis Comarcals de Banyoles, 1986-1987: 103-120.
Sala B. & Masini F. 2007. Late Pliocene and Pleistocene small mammal chronology in the Italian peninsula. Quaternary International, 160: 4-16.
Scheyer T.M., Sander P.M., Joyce W.G., Böhme W. & Witzel U. (2007) - A plywood structure in the shell of fossil and living soft-shelled turtles (Trionychidae) and its evolutionary implications. Organisms, Diversity and Evolution, 7: 136-144.
Schoch R.R., Poschmann M. & Kupfer A. (2015) - The salamandrid Chelotriton paradoxus from Enspel and Randeck Maars (Oligocene–Miocene, Germany). Palaeobiodiversity and Palaeoenvironments, 95: 77-86.
Scholtes S.J., Arntzen J.W., Ajduković M. & Ivanović A. (2022) - Variation in vertebrae shape across small‐bodied newts reveals functional and developmental constraints acting upon the trunk region. Journal of Anatomy, 240: 639-646.
Sesé C. (1989) - Micromamíferos del Mioceno, Plioceno y Pleistoceno de la cuenca de Guadix-Baxa (Granada). Trabajos sobre el Neógeno-Cuaternario, 11: 185-213.
Sinitsa M.V. (2018) - Phylogenetic position of Sinotamias and the early evolution of Marmotini (Rodentia, Sciuridae, Xerinae). Journal of Vertebrate Paleontology, 38: e1419251.
Sinitsa M.V. & Pogodina N.V. (2019) - The evolution of early Spermophilus in eastern Europe and the antiquity of the Old World ground squirrels. Acta Palaeontologica Polonica, 64: 643-667
Solé L., Macau C., Virgili C. & Llamas M.R. (1965) - Sobre los depósitos pliocénicos y cuaternarios del Bajo Ebro. Memorias y Comunicaciones del Instituto Jaime Almera CSIC, 1: 83-92.
Solé de Porta N. & Valle M.F. (1986) - Estudio palinológico del Plioceno de San Onofre (Tarragona). Estudios Geológicos, 42: 63-67.
Spagnesi M. & De Marinis A.M. (2002) - Mammiferi d’Italia. Quaderni di Conservazione della Natura, 14: 1-310.
Stefen C. (2005) - Description of the cranial morphology of the Early Miocene beaver Steneofiber castorinus. Neues Jahrbuch fur Geologie und Paläontologie Monatshefte, 2005: 577-596.
Stefen C. (2009) - Intraspecific variability of beaver teeth (Castoridae: Rodentia). Zoological Journal of the Linnean Society, 155: 926-936.
Stirton R.A. (1935) - A review of Tertiary beavers. University of California Publications, Bulletin of the Department of Geological Sciences, 23: 391-458.
Storch G. (1995) - The Neogene mammalian faunas of Ertemte and Harr Obo in Inner Mongolia (Nei Mongol), China. 11. Soricidae (Insectivora). Senckenbergiana Lethaea, 75: 221-251.
Suata-Alpaslan F. (2009) - Occitanomys (Rhodomys) vandami nov. sp. (Muridae, Mammalia) from the Early Pliocene/Early Ruscinian Fauna of Iğdeli (Turkey). The Open Geology Journal, 3: 64-69.
Szyndlar Z. (1984) - Fossil snakes from Poland. Acta Zoologica Cracoviensia, 28: 1-156.
Torres-Roig E., Piñero P., Agustí J., Bover P., Alcover J.A. (2019) - First evidence of endemic Murinae (Rodentia, Mammalia) in the early Pliocene of the Balearic Islands (western Mediterranean). Geological Magazine, 156: 1742-1750.
Van Dam J.A. (1997) - The small mammals from the Upper Miocene of the Teruel-Alfambra region (Spain): Paleobiology and paleoclimatic reconstructions. Geologica Ultraiectina, 156: 1-204.
Van Dam J.A. (2006) - Geographic and temporal patterns in the late Neogene (12-3 Ma) aridification of Europe. The use of small mammals as paleoprecipitation proxies. Paleogeography, Paleoclimatology, Paleoecology, 238: 190-218.
Van Dam J.A. & Utescher T. (2016) - Plant- and micromammal-based paleoprecipitation proxies: Comparing results of the Coexistence and Climate-Diversity Approach. Palaeogeography, Palaeoclimatology, Palaeoecology, 443: 18-33.
Van de Weerd A. (1976) - Rodent faunas of the Mio-Pliocene continental sediments of the Teruel-Alfambra region, Spain. Utrecht Micropaleontological Bulletin, Special Publication, 2: 1-217.
Van de Weerd A. (1979) - Early Ruscinian rodents and lagomorphs (Mammalia) from the lignites near Ptolemais (Macedonia, Greece). Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, 82: 127-170.
van den Hoek Ostende L.W. (2001) - A revised generic classification of the Galericini (Insectivora, Mammalia) with some remarks on their palaeobiogeography and phylogeny. Geobios, 34: 681-695.
Van der Made J. (2003) - Suoidea (Artiodactyla). In: Fortelius M., Kappelman J., Sen S. & Bernor R.L. (Eds) - Geology and Paleontology of the Miocene Sinap Formation, Turkey: 308-327. New York, Columbia University Press.
Van der Meulen A.J. & De Bruijn H. (1982) - The mammals from the Lower Miocene of Aliveri (Island of Evia, Greece). Part 2. The Gliridae. Proceedings Koninklijke Nederlandse Akademie van Wetenschappen B, 85: 485-524.
Venczel M. (2004) - Middle Miocene anurans from the Carpathian Basin. Palaeontographica Abteilung A, 271: 151-174.
Venczel M. & Hír J. (2013) - Amphibians and squamates from the Miocene of Felsőtárkány Basin, N-Hungary. Palaeontographica, Abteilung A, 300: 117-158.
Villa A., Delfino M., Luján À., Almécija S. & Alba D.M. (2019) - First record of Latonia gigantea (Anura, Alytidae) from the Iberian Peninsula. Historical Biology, 31: 371-382.
von Koenigswald W. & Reumer J.W.F. (2020) - The enamel microstructure of fossil and extant shrews (Soricidae and Heterosoricidae, Mammalia) and its taxonomical significance. Palaeontographica, Abteilung A, 316: 79-163.
Wessels W., Frieling J. & Fraaije R. (2011) - The oldest beaver from the Netherlands. Netherlands Journal of Geosciences, 90: 311-314.
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Copyright (c) 2024 PEDRO PIÑERO, JORDI AGUSTÍ, HUGUES-ALEXANDRE BLAIN, DAVID M. ALBA, MARC FURIÓ, CHIARA ANGELONE, ADRIANA LINARES-MARTÍN, RAFAEL MARQUINA-BLASCO, CASTO LABORDA-LÓPEZ, ALVARO ARASA-TULIESA, ISAAC CASANOVAS-VILAR
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Accepted 2024-07-12
Published 2024-09-10
