ORDOVICIAN (DARRIWILIAN–KATIAN) BRACHIOPODS FROM THE SOUTHEASTERN MARGIN OF AVALONIA (CONDROZ INLIER, BELGIUM)
DOI:
https://doi.org/10.54103/2039-4942/26428Keywords:
Brachiopoda; taxonomy; Ordovician; Avalonia; Belgium; palaeobiogeography.Abstract
A low-diversity, relatively well-preserved brachiopod fauna is documented in detail for the first time from the Sandbian–Katian (Upper Ordovician) siliciclastic rocks of the Oxhe Inlier (Belgium), i.e. a small Caledonian inlier included in the Condroz Inlier, whereas some preliminary data are also presented for the Darriwilian–Sandbian (Middle–Upper Ordovician) brachiopods of the central part of the Condroz Inlier (Huy and Vitrival-Bruyère formations), Belgium. The brachiopods from the Oxhe Inlier (Oxhe Formation) were collected from several localities in the Oxhe Inlier where they are associated with trilobites, ostracods, molluscs, echinoderms and machaeridians. The Oxhe Formation yielded one lingulate, one craniate, two strophomenates and three rhynchonellates, while the Huy Formation yielded two lingulates to which are added one strophomenate and two rhynchonellates from the Vitrival-Bruyère Formation (Sart-Bernard Member). These taxa are assigned with variable degrees of confidence to existing taxa or described under open nomenclature. An age range of Darriwilian to Katian is confirmed. Multivariate numerical analyses (Nonmetric Multidimensional Scaling together with Network Analysis) indicate close comparison with other Avalonian faunas. Moreover, the combination of taxa suggests habitats in deeper-water environments around the Avalonian continent.
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Bancroft B.B. (1928) - On the notational representation of the rib-system in Orthacea. Manchester Literary and Philosophical Society, Memoirs and Proceedings, 72: 53-90.
Bancroft B.B. (1929) - Some new genera and species of Strophomenacea from the Upper Ordovician of Shropshire. Memoirs and Proceedings of the Manchester Literary and Philosophical Society, 73: 33-65.
Bancroft B.B. (1945) - The brachiopod zonal indices of the stages Costonian to Onnian in Britain. Journal of Paleontology, 19: 181-252.
Barrande J. (1848) - Über die Brachiopoden der silurischen Schichten von Böhmen. Naturwissenschaftliche Abhandlungen, 2: 155-256.
Barrande J. (1879) - Système Silurien du centre de la Bohême. 1re partie. Recherches paléontologiques, Vol. 5, Classe de Mollusques, Ordre des Brachiopodes. Published by the Author, Prague and Paris, 226 pp.
Bastian M., Heymann S. & Jacomy M. (2009) - Gephi: an open source software for exploring and manipulating networks. Proceedings of the international AAAI conference on web and social media, 3(1): 361-362.
Bates D.E.B. (1968) - The Lower Palaeozoic brachiopod and trilobite faunas of Anglesey. Bulletin of the British Museum (Natural History), Geology, 16(4): 125-199.
Belanger I., Delaby S., Delcambre B., Ghysel P., Hennebert M., Laloux M., Marion J.-M., Mottequin B. & Pingot J.-L. (2012) - Redéfinition des unités structurales du front varisque utilisées dans le cadre de la nouvelle Carte géologique de Wallonie (Belgique). Geologica Belgica, 15(3): 169-175.
Benedetto J.L. (1995) - La fauna de braquiópodos de la Formación Las Plantas (Ordovícico tardío, Caradoc), Precordillera Argentina. Revista Española de Paleontología, 10(2): 239-258.
Béthune P. de (1954) - Carte géologique de Belgique. 1/500 000. Atlas de Belgique, planche 8. In: Fourmarier P. (Ed.), Prodrome d’une description géologique de la Belgique: Annexe 1. Société géologique de Belgique, Liège.
Boucot A.J. (1975) - Evolution and extinction rate controls. Elsevier, Amsterdam, 427 pp.
Bowdler-Hicks A., Ingham J.K. & Owen A.W. (2002) - The taxonomy and stratigraphical significance of the Anglo-Welsh Cryptolithinae (Trinucleidae, Trilobita). Palaeontology, 45: 1075-1105.
Candela Y. (2003) - Late Ordovician brachiopods from the Bardahessiagh Formation of Pomeroy, Co. Tyrone, Northern Ireland. Monograph of the Palaeontographical Society, 156(618): 1-95.
Candela Y. (2006) - Late Ordovician brachiopod faunas from Pomeroy, Northern Ireland: a paleoenvironmental synthesis. Transactions of the Royal Society of Edinburgh: Earth Sciences, 96: 317-325.
Candela Y. & Harper D.A.T. (2010) - Late Ordovician (Katian) brachiopods from the Southern Uplands of Scotland: Biogeographic patterns on the edge of Laurentia. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 100: 253-274.
Candela Y. & Mottequin B. (2022) - Tremadocian and Floian (Ordovician) linguliformean brachiopods from the Stavelot–Venn Massif (Avalonia; Belgium and Germany). Geologica Belgica, 22(1/2): 1-15.
Candela Y. & Mottequin B. (2023) - Middle and Upper Ordovician linguliformean and craniiformean brachiopods from the Brabant Massif, Belgium: infaunal giants, encrusting forms and durophagy. Geobios, 81: 101-119.
Candela Y., Marion J.-M., Servais T., Wang W., Wolvers M. & Mottequin B. (2021) - New linguliformean brachiopods from the lower Tremadocian (Ordovician) of the Brabant Massif, Belgium, with comments on contemporaneous faunas from the Stavelot–Venn Massif. Rivista Italiana di Paleontologia e Stratigrafia, 127(2): 383-395.
Cave R. & Dean W.T. (1959) - Four British Ordovician species of dalmanelloid brachiopod. Palaeontology, 1: 292-297.
Cluysenaar P.-G. & Lecrenier A. (1887) - Etude des fossiles siluriens de Huy et d’Ombret. Première communication. Bulletin du Cercle des Naturalistes hutois, 1887(2): 47-54.
Cocks L.R.M. (1978) - A review of British Lower Palaeozoic brachiopods, including a synoptic revision of Davidson’s monograph. Monographs of the Palaeontographical Society, 131(549): 2-228.
Cocks L.R.M. (2008) - A revised review of British Lower Palaeozoic brachiopods. Monograph of the Palaeontographical Society, 161(629): 1-276.
Cocks L.R.M. (2010) - Caradoc strophomenoid and plectambonitoid brachiopods from Wales and the Welsh Borderland. Palaeontology, 53: 1155-1200.
Cocks L.R.M. & Lockley M.G. (1981) - Reassessment of the Ordovician brachiopods form the Budleigh Salterton Pebble Bed, Devon. Bulletin of the British Museum (Natural History), Geology, 35(3): 111-124.
Cocks L.R.M. & Rong Jia-Yu (1989) - Classification and review of the brachiopod superfamily Plectambonitacea. Bulletin of the British Museum (Natural History), Geology, 45: 77-163.
Cocks L.R.M. & Torsvik T.H. (2002) - Earth geography from 500 to 400 million years ago: a faunal and palaeomagnetic review. Journal of the Geological Society, London, 159: 631-644.
Cocks L.R.M. & Torsvik T.H. (2021) - Ordovician palaeogeography and climate change. Gondwana Research, 100: 53-72.
Cooper G.A. (1956) - Chazyan and related brachiopods. Smithsonian Miscellaneous Collections, 127(I/II): 1-1245.
Dalman J.W. (1828) - Uppställning och Beskrifning af de i Sverige funne Terebratuliter. Kongliga Svenska Vetenskapsakademien, Handlingar for 1827: 85-155.
Davidson T. (1869) - A Monograph of the British Fossil Brachiopoda. Part 7, no. 3. The Silurian Brachiopoda. Monograph of the Palaeontographical Society, 22(96): 169-248.
Davidson T. (1870) - Notes on the Brachiopoda hitherto obtained from the ‘Pebble-bed’ of Budleigh Salterton, near Exmouth, in Devonshire. Quarterly Journal of the Geological Society London, 26: 70-90.
Davidson T. (1871) - A Monograph of the British Fossil Brachiopoda. Part 7, no. 4. The Silurian Brachiopoda. Monograph of the Palaeontographical Society, 24(108): 249-397.
Dean W.T. (1991) - Ordovician trilobites from the inlier at Le Petit Fond d’Oxhe, Belgium. Bulletin de l’Institut royal des Sciences naturelles de Belgique, Sciences de la Terre, 61: 135-155.
Denayer J. & Mottequin B. (2024) - The Lower Devonian lithostratigraphy of Belgium. Geologica Belgica, 27(3/4): 115-154. https://doi.org/10.20341/gb.2024.004
Destinez P. (1896) - Quelques nouveaux fossiles siluriens à Ombret. Annales de la Société géologique de Belgique, 23 (Bulletin): 118-119.
Dewalque G. (1894) - Un nouveau gîte de fossiles siluriens à Ombret. Annales de la Société géologique de Belgique, 21 (Bulletin): 80-81.
Forir H. (1899) - Compte rendu de la session extraordinaire de la Société géologique de Belgique, tenue à Huy, du 2 au 5 octobre 1897. Annales de la Société géologique de Belgique, 24 (pro 1897) (Bulletin): 149-203.
Fortey R.A., Harper D.A.T., Ingham J.K., Owen A.W. & Rushton A.W.A. (1995) - A revision of Ordovician series and stages from the historical type area. Geological Magazine, 132: 15-30.
Fortey R.A., Harper D.A.T., Ingham J.K., Owen A.W., Parkes, M.A., Rushton A.W.A. & Woodcock N. (2000) – Revised correlation of Ordovician rocks in the British Isles. Geological Society of London, London, 86 pp.
Gorjansky V.Iu. & Popov L.E. (1985) - Morfologiia, systematicheskoe polozhenie I proiskhozhdenie bezzamkovykh brakhiopods karbonatnoi rakovinoi [Morphology, systematic position and origin of the inarticulate brachiopods with calcareous shells]. Paleontologicheskii Zhurnal, 1985(3): 3-14 [in Russian].
Hall J. & Clarke J.M. (1892) - An introduction to the study of the genera of Palaeozoic Brachiopoda, Natural History of New York, Palaeontology, vol. 8, part 1. New York Geological Survey. Charles van Benthuysen & Sons, Albany, 367 pp.
Hammer Ø., Harper D.A.T. & Ryan P.D. (2001) - PAST: paleontological statistics software package for education and data analysis. Palaeontologia Electronica, 4. http://palaeo-electronica.org/2001-1/past/issue1_01.htm
Hansen J. (2008) - Upper Ordovician brachiopods from the Arnestad and Frognerkilen formations in the Oslo-Asker district, Norway. Palaeontos, 13: 1-99.
Hansen J. & Harper D.A.T. (2008) - The late Sandbian-earliest Katian (Ordovician) brachiopod immigration and its influence on the brachiopod fauna in the Oslo Region, Norway. Lethaia, 41: 25-35.
Harper D.A.T. (2000) - Dalmanellidina. In: Kaesler R.L. (Ed.) - Treatise on Invertebrate Paleontology, part H, Brachiopoda, Revised, Volume 3: Rhynchonelliformea (part): 782-844. The Geological Society of America and The University of Kansas. Boulder, Colorado and Lawrence, Kansas.
Harper D.A.T., Mitchell W.I., Owen A.W. & Romano M. (1985) - Upper Ordovician brachiopods and trilobites from the Clashford House Formation, near Herbertstown, Co. Meath, Ireland. Bulletin of the British Museum (Natural History), Geology Series, 38(5): 287-308.
Harper D.A.T., Parkes M.A. & Zhan Ren-Bin (2017) - Late Ordovician deep-water brachiopod fauna from Raheen, Waterford Harbour, Ireland. Irish Journal of Earth Sciences, 35: 1-18.
Harper D.A.T. & Wright A.D. (1996) - Brachiopods: 63–94. In: Harper D.A.T. & Owen A.W. (Eds) - Fossils of the Upper Ordovician. Palaeontological Association Field Guides to Fossils, 7: 63-94. The Palaeontological Association, London.
Havlíček V. (1950) - Ramenonožci českého ordoviku. Rozpravy Ústředního ústavu geologického, 13: 1-72.
Havlíček V. (1951) - The Ordovician Brachiopoda from Bohemia. Rozpravy Ústředního ústavu geologického, 12: 1-135.
Havlíček V. (1977) - Brachiopods of the order Orthida in Czechoslovakia. Rozpravy Ústředního ústavu geologického, 44: 1-327.
Havliček V. (1980) - Inarticulate brachiopods in the Lower Ordovician of the Montagne Noire. (South France). Mémoire de la Société d’Études scientifiques de l’Aude, 1980: 3-11.
Havliček V. (1982) - Lingulacea, Paterinacea, and Siphonotretacea (Brachiopoda) in the Lower Ordovician sequence of Bohemia. Sbornik geologickych věd, Paleontologie, 25: 9-82.
Havlíček V. & Branisa L. (1980) - Ordovician brachiopods of Bolivia (Succession of assemblages, climate control, affinity to Anglo-French and Bohemian provinces). Rozpravy Československé akademie věd, 90: 1-54.
Herbosch A. & Verniers J. (2014) - Stratigraphy of the Lower Palaeozoic of the Brabant Massif, Belgium. Part II: The Middle Ordovician to lowest Silurian of the Rebecq Group. Geologica Belgica, 17: 115-136.
Hints L. (1975) - Ordovician brachiopods: Enteletacea of the East Baltic area. Eesti NSV Teaduste Akadeemia Geoloogia, Tallinn, 117 pp. [in Russian].
Hurst J.M. (1978a) - A phenetic strategy model for dalmanellid brachiopods. Palaeontology, 21(3): 535-554.
Hurst J.M. (1978b) - Morphological variation in some Lower Palaeozoic dalmanellid brachiopods. Paleontologicheskii Zhurnal, 52: 246-256.
Hurst J.M. (1979a) - The stratigraphy and brachiopods of the upper part of the type Caradoc of south Salop. Bulletin of the British Museum (Natural History), Geology, 32: 183-304.
Hurst J.M. (1979b) - The environment of deposition of the Caradoc Alternata Limestone and contiguous deposits of Salop. Geological Journal, 14(1): 15-40.
Jones C.R. & Siveter D.J. (1983) - On Harperopsis scripta (Harper). Stereo-atlas of ostracod shells, 10: 5-12.
Jones O.T. (1928) - Plectambonites and some allied genera. Memoirs of the Geological Survey of Great Britain, Palaeontology, 12(5): 367-527.
Lecrenier A. (1887) - Sur des fossiles siluriens de Huy. Annales de la Société géologique de Belgique, 14 (Bulletin): 182-183.
Lefebvre B., Javier Álvaro J., Casas J.M., Ghienne J.-F., Herbosch A., Loi A., Monceret E., Verniers J., Vidal M., Vizcaïno D. & Servais T. (2023) - The Ordovician of France and neighbouring areas of Belgium and Germany. Geological Society, London, Special Publications, 532: 375-408.
Liljeroth M., Harper D.A.T., Carlisle H. & Nielsen A.T. (2017) - Ordovician rhynchonelliformean brachiopods from Co. Waterford, SE Ireland: Palaeobiogeography of the Leinster Terrane. Fossils and Strata, 62: 1-164.
Lockley M.G. (1980) - The Caradoc faunal associations of the area between Bala and Dinas Mawddy, north Wales. Bulletin of the British Museum (Natural History), Geology, 33: 165-235.
Lockley M.G. (1983) - A review of brachiopod dominated palaeocommunities from the type Ordovician. Palaeontology, 26(1): 111-145.
Lohest M. & Forir H. (1900) - Quelques découvertes intéressantes faites pendant les excursions du cours de géologique de l’Université de Liége. Annales de la Société géologique de Belgique, 27 (Bulletin): 161-163.
Maillieux E. (1926) - Remarques sur l’Ordovicien de la Belgique. Bulletin de la Société belge de Géologie, de Paléontologie et d’Hydrologie, 27 (Procès-verbaux): 67-85.
Maillieux E. (1933) - Terrains, roches et fossiles de la Belgique. Deuxième édition revue et corrigée. Patrimoine du Musée royal d’Histoire naturelle de Belgique, Bruxelles, 217 pp.
Maillieux E. (1938a) - Observations nouvelles sur l’Ordovicien de la Belgique. Bulletin de la Société belge de Géologie, de Paléontologie et d’Hydrologie, 48 (Procès-verbaux): 22-23.
Maillieux E. (1938b) - Compte rendu de l’excursion de la Société belge de Géologie à Sart-Bernard et à Naninne, le 30 juin 1938. Bulletin de la Société belge de Géologie, de Paléontologie et d’Hydrologie, 48 (Procès-verbaux): 338-341.
Maillieux E. (1939) - L’Ordovicien de Sart-Bernard. Mémoires du Musée royal d’Histoire naturelle de Belgique, 86: 1-59.
Malaise C. (1873) - Description du terrain silurien du centre de la Belgique. Mémoires couronnés et mémoires des savants étrangers publiés par l’Académie royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 37: 1-122.
Malaise C. (1887a) - Observations sur la lettre précédente. Annales de la Société géologique de Belgique, 14 (Bulletin): 183.
Malaise C. (1887b) - Observations sur quelques graptolithes de la bande silurienne de Sambre-et-Meuse. Annales de la Société géologique de Belgique, 14 (Bulletin): 183-184.
Malaise C. (1888a) - Découverte de la faune de la base du Silurien en Belgique. Bulletin de l’Académie royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 3e série, 15: 365-367.
Malaise C. (1888b) - Les schistes siluriens de Huy et leur signification géologique. Annales de la Société géologique de Belgique, 15 (Bulletin): 39-44.
Malaise C. (1888c) - A propos des schistes siluriens de Huy. Annales de la Société royale malacologique de Belgique, 23 (Bulletin des séances): 10-13.
Malaise C. (1890) - Sur les graptolithes de Belgique. Bulletin de l’Académie royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 3e série, 20: 440-452.
Malaise C. (1894a) - Sur l’aspect llandeilien du massif d’Oxhe. Annales de la Société géologique de Belgique, 21 (Bulletin): 108-110.
Malaise C. (1894b) - Sur la découverte de Beyrichia dans le Silurien belge. Annales de la Société géologique de Belgique, 21 (Bulletin): 95.
Malaise C. (1899) - Espèces nouvelles de la bande silurienne de Sambre-et-Meuse. Annales de la Société géologique de Belgique, 24 (pro 1897) (Mémoires): 257-261.
Malaise C. (1900a) - Etat actuel de nos connaissances sur le Silurien de la Belgique (à continuer). Annales de la Société géologique de Belgique, 25bis: 179-216.
Malaise C. (1900b) - A propos du massif silurien du Fond d’Oxhe. Annales de la Société géologique de Belgique, 27 (Bulletin): 167-168.
Malaise C. (1901a) - Etat actuel de nos connaissances sur le Silurien de la Belgique (fin). Annales de la Société géologique de Belgique, 25bis: 217-221.
Malaise C. (1901b) - Sur le Silurien de Belgique. In: Congrès géologique international. Comptes rendus de la 8e session en France, tenu à Paris du 16 au 20 août 1900. Premier fascicule: pp. 561-571. Le Bigot Frères, Paris.
Malaise C. (1911) - Sur l’évolution de l’Echelle stratigraphique du Siluro-cambrien de Belgique. Annales de la Société géologique de Belgique, 38: BB7-BB28.
Martin F. (1969) - Les Acritraches de l’Ordovicien et du Silurien Belge. Détermination et valeur stratigraphique. Mémoires de l’Institut royal des Sciences naturelles de Belgique, 160 (pro 1968): 1-175.
M’Coy F. (1846) - A synopsis of the Silurian fossils of Ireland collected from the several districts by Richard Griffith F.G.S. Privately published, Dublin, 72 pp.
Menke C.T. (1828) - Synopsis methodica molluscorum generum omnium et specierum earum quae in Museo Menkeano adservantur. G. Uslar, Pyrmonti, 91 pp.
Mergl M. (2002) - Linguliformean and craniiformean brachiopods of the Ordovician (Třenice to Dobrotiva Formations) of the Barrandian, Bohemia. Acta Musei Nationalis Pragae, Series B, Natural History, 58(1-2): 1-82.
Michot P. (1934) - Le massif ordovicien du Fond d’Oxhe. Annales de la Société géologique de Belgique, 57(4): B59-B79.
Michot P. (1954) - Le Silurien. In: Fourmarier, P. (Ed.) - Prodrome d’une description géologique de la Belgique: 39-82. Société géologique de Belgique, Liège.
Michot P. (1957) - Assise d’Oxhe (ou Fond d’Oxhe). In: Waterlot G. (Ed.) - Fascicule 4a: France, Belgique, Pays-Bas, Luxembourg. Fascicule 4a1: Antécambrien et Paléozoïque inférieur. In: Pruvost P. (Ed.) - Lexique Stratigraphique International. Volume 1 (Europe): 316-317. C.N.R.S, Paris.
Michot P. (1969) - La faille d’Ombret. Annales de la Société géologique de Belgique, 92(2): 243-254.
Moore R.C. (1952) - Brachiopoda. In: Moore R.C., Lalicker C.G. & Fischer A.G. - Invertebrate Fossils: 197-267. McGraw-Hill, New York.
Mortier J., Vanmeirhaeghe J., Harper D.A.T., Štorch P., Zalasiewicz J., Van Den Haute P., Deckers J., Mestdagh T., Pille T. & Verniers J. (2024) - Stratigraphy, biostratigraphy, and chitinozoans of the uppermost Ordovician and Silurian of the Condroz Inlier. Memoirs of the Geological Survey of Belgium, 65: 1-239.
Mottequin B. (2021) - Earth science collections of the Centre Grégoire Fournier (Maredsous) with comments on Middle Devonian–Carboniferous brachiopods and trilobites from southern Belgium. Geologica Belgica, 24(1-2): 33-68.
Mottequin B., Marion J.-M. & Delcambre B. (2021) - Carte géologique de Wallonie: Huy – Nandrin 48/3-4. 1/25 000. Service public de Wallonie, Agriculture, Ressources naturelles et Environnement, Namur, avec une notice explicative de 115 p.
Öpik A.A. (1930) - Brachiopoda Protremata der Estländischen Ordovizischen Kukruse-Stufe. Universitatis Tartuensis (Dorpatensis) Acta et Commentationes, series A, 17: 1-262.
Öpik A.A. (1934) - Über die Klitamboniten. Tartu Ulikooli Geoloogia-Instituudi Toimestuesed Acta et Commentationes Universitatis Tartuensis, 39: 1-239.
Owens R.M. & Servais T. (2007) - The Ordovician of the Condroz Inlier, Belgium: Trilobites from the southeastern margin of Avalonia. Palaeogeography, Palaeoclimatology, Palaeoecology, 245: 272-294.
Parkes M.A. (1994) - The brachiopods of the Duncannon Group (Middle-Upper Ordovician) of southeast Ireland. Bulletin of the Natural History Museum (Geology) 50(2): 105-174.
Paškevičius J. (1994) - Baltijos Respublikų Geologija [=The Geology of the Baltic Republics]. Vastybinis leidybos centras, Vilnius, 447 pp. [in Lithuanian].
Pickerill R.K. & Brenchley P.J. (1979) - Caradoc marine benthic communities of the South Berwyn Hills, North Wales. Palaeontology, 22(1): 229-264.
Pohl A., Nardin E., Vandenbroucke T.R.A. & Donnadieu Y. (2016) - High dependence of Ordovician ocean surface circulation on atmospheric CO2 levels. Palaeogeography, Palaeoclimatology, Palaeoecology, 458: 39-51.
Popov L.E., Bassett M.G., Holmer L.E. & Laurie J.R. (1993) - Phylogenetic analysis of higher taxa of Brachiopoda. Lethaia, 26: 1-5.
Popov L.E. & Pushkin V.I. (1986) – Ordovikskie bezzamkovye brahiopody ûžnoj Pribaltiki [Ордовикские беззамковые брахиоподы южной Прибалтики]. Novyie i maloizvestnye vidy iskopaemykh životnyh i rastenij Belorussii: 11-22. Nauka i Tekhnika, Minsk.
Potter A.W. & Boucot A.J. (1992) - Middle and late Ordovician brachiopod benthic assemblages of North America. In: Webby B.D. & Laurie J.R. (Eds) - Global perspectives on Ordovician Geology: 307-323. Balkema, Rotterdam.
Rasmussen C.M.Ø. (2011) - Final destination, first discovered: the tale of Oanduporella Hints, 1975. In: Gutiérrez-Marco J.C., Rábano I. & García-Bellido D. (Eds) - Ordovician of the World: 447-454. Cuadernos del Museo Geominero, 14. Instituto Geológico y Minero de España, Madrid.
Rasmussen C.M.Ø., Harper D.A.T. & Blodgett R.B. (2012) - Late Ordovician brachiopods from west-central Alaska: systematics, ecology and palaeobiogeography. Fossils and Strata, 58:1-103
Reed F.R.C. (1910) - New fossils from the Dufton Shales. Geological Magazine, 7(5): 211-220, 294-299.
Rong J.-y. & Cocks L.R.M. (1994) - True Strophomena and a revision of the classification and evolution of strophomenoid and ‘strophodontid’ brachiopods. Palaeontology, 37(3): 651-694.
Salter J.W. (1852) - A systematic description of Palaeozoic fossils in the Geological Museum of the University of Cambridge. Appendix A. Description of a few species from Wales and Westmoreland referred to in the foregoing work [Appendix to Sedgwick A. & McCoy F. 1851], i-viii.
Salter J.W. & Aveline W.T. (1854) - On the “Caradoc Sandstone” of Shropshire. Quarterly Journal of the Geological Society, London, 10: 62-75.
Schallreuter R., Verniers J. & De Geest P. (2000) - An Ordovician ostracod from Belgium. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte, 2000(9): 570-576.
Schuchert C. (1893) - Classification of the Brachiopoda. American Geologist, 11: 141-167.
Schuchert C. (1913) - Class 2. Brachiopoda. In: von Zittel K. A. (Ed.) - Text-book of Palaeontology, vol. 1, part 1, 2nd edition, translated and edited by C. R. Eastman: 355-420. MacMillan and Co. Ltd., London.
Schuchert C. & Cooper G.A. (1932) - Brachiopod genera of the suborders Orthoidea and Pentameroidea. Memoirs of the Peabody Museum of Natural History, 4(1): 1-270.
Sedgwick A. & M’Coy F. (1852) - A synopsis of the classification of the British Palaeozoic rocks, with a systematic description of the British Palaeozoic fossils in the geological Museum of the University of Cambridge. London and Cambridge, xcviii + 661 pp., 25 pls.
Servais T. & Maletz J. (1992) - Lower Llanvirn (Ordovician) graptolites and acritarchs from the ‘Assise de Huy’, Bande de Sambre-et-Meuse, Belgium. Annales de la Société géologique de Belgique, 115(1): 265-285.
Sheehan P.M. (1987) - Late Ordovician (Ashgillian) brachiopods from the region of the Sambre and Meuse Rivers, Belgium. Bulletin de l’Institut royal des Sciences naturelles de Belgique, Sciences de la Terre, 57: 5-81.
Sinclair G.W. (1946) - Bancroftina, a new brachiopod name. Journal of Paleontology, 20: 295.
Siveter D.J. (1984) - Ecology of Silurian ostracods. Special Papers in Palaeontology, 32: 71-85.
Siveter D.J. (2009) - Ordovician. In: Whittaker J.E. & Hart M.B. (Eds) - Ostracods in British Stratigraphy: 15-44. The Geological Society for the Micropalaeontological Society, Special Publications, London.
Sowerby J. de C. (1839) - Mollusca and Conchifers. In: Murchison R.I. - The Silurian System, Part 2. Organic remains: 577-768. J. Murray, London.
Suárez-Soruco R. (1992) - El Paleozoico Inferior de Bolivia y Perú. In: Gutiérrez-Marco J.C., Saavedra J. & Rábano I. (Eds.) - Paleozoico Inferior de Ibero-América: 225-239. Universidad de Extremadura, Mérida.
Vanmeirhaeghe J. (2006) - The evolution of the Condroz-Brabant Basin from Middle Ordovician to Llandovery: lithostratigraphical and chitinozoan biostratigraphical approach. Unpublished Ph.D. thesis, Ghent University, Belgium.
Vanmeirhaeghe J. (2007a) - Chitinozoans of the lower Llanvirn Huy and the middle Caradoc Sart-Bernard formations (Middle to Upper Ordovician): implications for the stratigraphy of the Condroz Inlier (Belgium). Carnets de Géologie/Notebooks on Geology, M01/10: 59-67.
Vanmeirhaeghe J. (2007b) - Review of the lithostratigraphy of the Middle Ordovician to Llandovery of the Condroz Inlier (Belgium) by chitinozoan biostratigraphy: does the sedimentary succession record the Ardennian deformation? Geologica Belgica, 10: 162-164.
Verniers J., Herbosch A., Vanguestaine M., Geukens F., Delcambre B., Pingot J.-L., Belanger I., Hennebert M., Debacker T., Sintubin M. & De Vos W. (2002) - Cambrian-Ordovician-Silurian lithostratigraphic units (Belgium). Geologica Belgica, 4 (pro 2001): 5-38.
Waagen W.H. (1884) - Salt Range fossils, vol. 1, part 4. Productus Limestone fossils, Brachiopoda. Memoirs of the Geological Survey of India, Palaeontologia Indica, series 13, 3-4: 547-728.
Waagen W.H. (1885) - Salt Range fossils, vol. 1, part 4. Productus Limestone fossils, Brachiopoda. Memoirs of the Geological Survey of India, Palaeontologia Indica, series 13, 5: 729-770.
Waisfeld B.G. & Henry J.L. (2003) - Huemacaspis (Trilobita, Kerfornellinae) from the Late Ordovician of the Argentine Cordillera Oriental. Geobios, 36: 491-499.
Whittington H.B. (1938) - New Caradocian brachiopods from the Berwyn Hills, North Wales. Annals and Magazine of Natural History, series 11, 2: 241-259.
Williams A. (1962) - The Barr and Lower Ardmillan Series (Caradoc) of the Girvan District, South-west Ayrshire, with descriptions of the Brachiopoda. Geological Society of London Memoir, 3: 1-267.
Williams A. (1963) - The Caradocian brachiopod faunas of the Bala District, Merionethshire. Bulletin of the British Museum (Natural History), Geology, 8(7): 327-471.
Williams A. (1974) - Ordovician Brachiopoda from the Shelve District, Shropshire. Bulletin of the British Museum (Natural History), Geology, Supplement, 11: 1-163.
Williams A., Carlson S.J., Brunton C.H.C., Holmer L.E. & Popov L.E. (1996) - A supra-ordinal classification of the Brachiopoda. Philosophical Transactions of the Royal Society of London, series B, 351: 1171-1193.
Williams A. & Wright A.D. (1963) - The classification of the “Orthis testudinaria Dalman” group of brachiopods. Journal of Paleontology, 37: 1-32.
Wright A.D. (1964) - The fauna of the Portrane Limestone II. Bulletin of the British Museum (Natural History), Geology, 9: 137-256.
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