CARINTHIASUCHUS KANDUTSCHI, A NEW ARCHOSAUR (REPTILIA: DIAPSIDA) FROM THE TRIASSIC OF AUSTRIA
DOI:
https://doi.org/10.54103/2039-4942/29890Abstract
A partial skeleton of a small reptile with a distinctive maxilla, dentition, caudal vertebrae and osteoderms
from the marine Partnach Formation (Ladinian-earliest Carnian) near Jadersdorf (Gitschtal, Hermagor) in the Gailtal Alps
of Carinthia (Austria), represents a new archosaur taxon Carinthiasuchus kandutschi gen. n. sp. n.. It is diagnosed by autapomorphies
in the anterior process and subnarial foramen of the maxilla, accessory articular processes in the neural arches of
mid-distal caudal vertebrae, and shape and arrangement of the caudal osteoderms. It also shows a unique combination of
features regarding the extent of the antorbital fossa in the maxilla and the morphology of its ventral edge, dentary shape,
dentition, caudal vertebrae elongation and morphology. A segment of 16 mid-distal caudal vertebrae is one of the better articulated
tail segments ever found in Triassic archosaurs, showing the arrangement of the dorsal osteoderms and the presence
of accessory processes in the neural arches stiffening the posterior termination of the tail. Phylogenetic analysis recovered
Carinthiasuchus kandutschi as a pseudosuchian in a basal position within the Poposauroidea. Absence of swimming adaptation
in the tail and the presence of terrestrial plant remains in the same outcrop suggest terrestrial habits for this archosaur reptile.
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References
Bachmayer H. & Warch A (1959) - Ein bemerkenswerter Fischfund in den Partnachschichten (Trias) südlich von Feistritz an der Drau (Kärnten). Carinthia II, 69: 42-43.
Bechstädt T. (1978) - Faziesanalyse permischer und triadischer Sedimente des Drauzuges als Hinweis auf eine großräumige Lateralverschiebung innerhalb des Ostalpins. Jahrbuch der Geologischen Bundes-Anstalt, 121: 1-121.
Bonaparte J. F. (1981) - Descripcion de “Fasolasuchus tenax” y su significado en la sistematica y evolución de los thecodontia. Revista del Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, III (2): 55-101.
Butler R.J., Nesbitt S.J., Charig A.J., Gower D.J. & Barrett P.M. (2017) - Mandasuchus tanyauchen, gen. et sp. nov., a pseudosuchian archosaur from the Manda Beds (?Middle Triassic) of Tanzania, Journal of Vertebrate Paleontology, 37 (suppl.1): 96-121.
Charig A.J., Krebs B., Sues H.-D. & Westphal F. (1976) - Thecodontia. In: Kuhn O. (Ed.) - Handbuch der Paläoherpetologie, 13: 40-98. G. Fischer Verlag. Stuttgart.
Colbert E.H. & Mook C.C. (1951) - The ancestral crocodilian Protosuchus. Bulletin of the American Museum of Natural History, 97:143-182.
Cope E.D. (1869) - Synopsis of the extinct Batrachia, Reptilia and Aves of North America. Transactions of the American Philosophical Society, 14:1-252.
Crush P.J. (1984) - A late Upper Triassic sphenosuchid crocodilian from Wales. Palaeontology, 27: 131-157.
Dalla Vecchia F.M. (2006) - Resti di Tanystropheus, saurotterigi e “rauisuchi” (Reptilia) nel Triassico medio della Val Aupa (Moggio Udinese, Udine). Gortania 27(2005): 25-48.
Dalla Vecchia F.M. (2008) - Vertebrati fossili del Friuli - 450 milioni di anni di evoluzione. Pubblicazioni del Museo Friulano di Storia Naturale, n. 50, 304 pp., 279 figs, Udine.
Dalla Vecchia F.M. (2014) - Gli pterosauri triassici. Pubblicazione del Museo Friulano di Storia Naturale n. 54, Museo Friulano di Storia Naturale, Udine, 319 pp.
Dalla Vecchia F.M. (2021) - Heteropelta boboi n. gen., n. sp. an armored archosauriform (Reptilia: Archosauromorpha) from the Middle Triassic of Italy. PeerJ, 9: e12468 (24 pp.) doi 10.7717/peerj.12468
Dalla Vecchia F.M. & Avanzini M. (2002) - New scattered remains of Triassic reptiles from northeastern Italy. Bollettino della Società Paleontologica Italiana, 41(2/3): 215-235.
Dalla Vecchia F.M. & Simonetto L. (2018) - Osteological remains of reptiles from the Friuli region (NE Italy) in the palaeontological collections of the Museo Friulano di Storia Naturale. Gortania - Geologia, Paleontologia, Paletnologia, 39 (2017): 27-70.
Dalla Vecchia F.M., Wild R., Hopf H. & Reitner J.A. (2002) - A crested rhamphorhynchoid pterosaur from the Late Triassic of Austria. Journal of Vertebrate Paleontology, 22:195-198.
Dawley R.M., Zawiske J.M. & Cosgriff J.W. (1979) - A rauisuchid thecodont from the Upper Triassic Popo Agie formation of Wyoming. Journal of Paleontology, 53:1428-1431.
Desojo J.B. & Rauhut O.W.M. (2024) - Reassessment of the enigmatic “Prestosuchus” loricatus (Archosauria: Pseudosuchia) from the Middle-Late Triassic of southern Brazil. The Anatomical Record, 307(4): 974-1000.
Ezcurra M.D. (2016) - The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms. PeerJ, 4: e1778.
França M.A.G., Ferigolo J. & Langer M.C. (2011) - Associated skeletons of a new middle Triassic “Rauisuchia” from Brazil. Naturwissenschaften, 98: 389-395.
Goloboff P. & Morales M. (2023) - TNT version 1.6, with a graphical interface for MacOs and Linux, including new routines in parallel. Cladistics, 39(2): 144-153.
Gower D.J. (1999) - The cranial and mandibular osteology of a new rauisuchian archosaur from the Middle Triassic of southern Germany. Stuttgarter Beiträge zur Naturkunde B (Geologie und Palaeontologie), 280: 1-49.
Gower D.J. & Schoch R. (2009) - Postcranial anatomy of the rauisuchian archosaur Batrachotomus kupferzellensis. Journal of Vertebrate Paleontology, 29: 103-122.
Harris J.D. & Downs A. (2002) - A drepanosaurid pectoral girdle from the Ghost Ranch (Whitaker) Coelophysis Quarry (Chinle Group, Rock Point Formation, Rhaetian), New Mexico. Journal of Vertebrate Paleontology, 22(1): 70-75.
Irmis R.B., Nesbitt S.J. & Sues H.-D. (2013) - Early Crocodylomorpha. In: Nesbitt S.J., Desojo J.B. & Irmis R.B. (Eds.) - Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin. Geological Society of London, Special Publication, 379: 275-302.
Klein N., Furrer H., Dojen C., Lukeneder A. & Scheyer T.M. (2025) - A new nothosaurid taxon from the Middle Triassic of Carinthia, Austria. Austrian Journal of Earth Sciences, 118(1):141-155.
Krebs B. (1965) - Die Triasfauna der Tessiner Kalkalpen. XIX. Ticinosuchus ferox, nov. gen. nov. sp. Ein neuer Pseudosuchier aus der Trias des Monte San Giorgio. Schweizerische Paläontologische Abhandlungen, 81: 1-140.
Laurenti J.N. (1768) - Classis Reptilium. Specimen medicum, exhibens synopsis Reptilium emendatum, cum experimentis circa venena et antidote Reptilium Austriacorum. Vienna: typis Johannes Thomae de Trattnern.
Lautenschlager S. & Desojo J.B. (2011) - Reassessment of the Middle Triassic rauisuchian archosaurs Ticinosuchus ferox and Stagonosuchus nyassicus. Paläontologische Zeitschrift, 85: 357-381.
Lautenschlager S. & Rauhut O.W.M. (2014) - Osteology of Rauisuchus tiradentes from the Late Triassic (Carnian) Santa Maria Formation of Brazil, and its implications for rauisuchid anatomy and phylogeny. Zoological Journal of the Linnean Society,173(1): 55-91.
Li C., Wu X.-C., Cheng Y.-N., Sato T. & Wang L.-T. (2006) - An unusual archosaurian from the marine Triassic of China. Naturwissenschaften, 93: 200-206.
Li C., Wu X.-C., Zhao L.-J., Nesbitt S.J., Stocker M. R. & Wang L.-T. (2016) - A new armored archosauriform (Diapsida: Archosauromorpha) from the marine Middle Triassic of China, with implications for the diverse life styles of archosauriforms prior to the diversification of Archosauria. The Science of Nature, 103: 95 (23 pp.).
Mastrantonio B.M., Baczko M.B. von, Desojo J.B. & Schultz C.L. (2019) - The skull anatomy and cranial endocast of the pseudosuchid archosaur Prestosuchus chiniquensis from the Triassic of Brazil. Acta Palaeontologica Polonica, 64(1): 171-198.
Modesto S.P. & Anderson J.S. (2004) - The phylogenetic definition of Reptilia. Systematic Biology, 53: 815-821.
Nesbitt S.J. (2005) - The osteology of the Middle Triassic pseudosuchian archosaur Arizonasaurus babbitti. Historical Biology. 17: 19-47.
Nesbitt S.J. (2011) - The early evolution of Archosauria: relationships and the origin of major clades. Bulletin of the American Museum of Natural History, 352: 1-292.
Nesbitt S.J. & Chatterjee S. (2024) - The osteology of Shuvosaurus inexpectatus, a shuvosaurid pseudosuchian from the Upper Triassic Post Quarry, Dockum Group of Texas, USA. The Anatomical Record, 307: 1175-1238.
Nesbitt S.J., Desojo J.B., Irmis R.B. (eds.) (2013) - Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin. Geological Society of London, Special Publication 379, 601 pp.
Nesbitt S.J., Liu J. & Li C. (2011) - A sail-backed suchian from the Heshanggou Formation (Early Triassic: Olenekian) of China. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 101: 271-284.
Nesbitt S.J., Zawiskie J.M. & Dawley R.M. (2020) - The osteology and phylogenetic position of the loricatan (Archosauria: Pseudosuchia) Heptasuchus clarki, from the ?Mid-Upper Triassic, southeastern Big Horn Mountains, Central Wyoming (USA). PeerJ, 8: e10101 (pp. 56). Doi 10.7717/peerj.10101
Nesbitt S.J., Stocker M.R., Small B.J. & Downs A. (2009) -The osteology and relationships of Vancleavea campi (Reptilia: Archosauriformes). Zoological Journal of the Linnean Society of London,157: 814-864.
Nesbitt S.J., Patellos E., Kammerer C.F., Ranivoharimanana L., Wyss A.R., Flynn J.J. (2023) - The earliest-diverging avemetatarsalian: a new osteoderm-bearing taxon from the Triassic (?Earliest Late Triassic) of Madagascar and the composition of avemetatarsalian assemblages prior to the radiation of dinosaurs. Zoological Journal of the Linnean Society, 199: 327-353.
Nicosia U., Avanzini M., Barbera C., Conti M.A., Dalla Vecchia F.M. and other 15 coauthors in alphabetic order (2005) - I vertebrati continentali del Paleozoico e Mesozoico. In: Bonfiglio L. (ed.), Paleontologia dei Vertebrati in Italia, Memorie del Museo Civico di Storia Naturale di Verona (serie 2, Sez. Scienze della Terra), 6: 41-66.
Nopcsa F. (1923) - Die Familien der Reptilien. Fortschritte der Geologie und Paläontologie, 2: 1-210.
Parker W.G., Nesbitt S.J., Irmis R.B., Martz J.W., Marsh A.D., Brown M.A., Stocker M.R. & Werning S. (2022) - Osteology and relationships of Revueltosaurus callenderi (Archosauria: Suchia) from the Upper Triassic (Norian) Chinle Formation of Petrified Forest National Park, Arizona, United States. The Anatomical Record, 305(10): 2353-2414.
Parrish J.M. (1993) - Phylogeny of the Crocodylotarsi, with reference to archosaurian and crurotarsan monophyly. Journal of Vertebrate Paleontology, 13: 287-308.
Perner T. (2018) - A new interesting archosaur from the Ladinian (Middle Triassic) of the Dolomites (Northern Italy). Preliminary report. In: Perner T. & Wachtler M. (eds), Some new and exciting Triassic Archosauria from the Dolomites (Northern Italy). Dolomythos Museum - Oregon Institute of Geological Research. Pp. 1-8.
Petti F.M., Bernardi M., Kustatscher E., Renesto S. & Avanzini M. (2013) - Diversity of continental tetrapods and plants in the Triassic of the Southern Alps: ichnological, paleozoological, and paleobotanical evidence. New Mexico Museum of Natural History and Science, Bulletin, 61: 458-484.
Piller W.E. Egger H., Erhart C.W., Gross M., Harzhauser M., Hubmann B., van Husen D., Krenmayr H.-G., Krystyn L., Lein R., Lukeneder A., Mandl G., Rögl F., Roetzl R., Rupp C., Schnabel W., Schönlaub H. P., Summesberger H., Wagreich M. & Wessley G. (2004) - Die stratigraphische Tabelle von Österreich 2004 (sedimentäre Schichtfolgen).
Renesto S., Confortini F., Gozzi E., Malzanni M. & Paganoni A. (2003) - A possible rauisuchid (Reptilia, Archosauria) tooth from the Carnian (Late Triassic) of Lombardy (Italy). Rivista del Museo Civico di Scienze Naturali “E. Caffi” Bergamo, 22: 109-114.
Rieppel O. (2019) - Mesozoic Sea Dragons: Triassic Marine Life from the Ancient Tropical Lagoon of Monte San Giorgio. Indiana University Press, Bloomington & Indianapolis, 256 pp.
Röhl H.J., Schmid-Röhl A., Furrer H., Frimmel A., Oschmann W. & Schwark L. (2001) - Microfacies, geochemistry and palaeoecology of the Middle Triassic Grenzbitumenzone from Monte San Giorgio (Canton Ticino, Switzerland). Geologia Insubrica, 6 (1): 1-13.
Schoch R. (2011) - New archosauriform remains from the German Lower Keuper. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, 260/1: 87-100.
Schönlaub H.P., Bär R., Bekmann U., Boekelmann K., Buttersack E., Fenninger A., Geiger A., Grossman J., Hauser Ch., Heinisch H., Van Husen D., Kahler F., Köhler M., Müller P.J., Niedermayr E., Niedermayr G., Pölsler P., Prey S., Suttner E., Überhorst S., Venturini C., Warch A. & Wilke M. (1987) - 198 - Weissbriach. Geologische Karte der Republik Österreich 1: 50.000, Geologische Bundesanstalt, Wien.
Scheyer T.M. & Desojo J.B. (2011) - Palaeohistology and external microanatomy of rauisuchian osteoderms (Archosauria: Pseudosuchia). Palaeontology, 54(6): 1289-1302.
Schönlaub H.P., Auferbauer H., Deutsch A., Heinisch H., Heinz H., Herzog U., Van Husen D., Lieberman H.M., Skala W.D. & Warch A. (1989) - 199 - Hermagor. Geologische Karte der Republik Österreich 1: 50.000, Geologische Bundesanstalt, Wien.
Sieber R. (1955) - Ein bemerkenswerter Fischfund aus der Mitteltrias Kärntens. Carinthia II, 65: 91-96.
Sill W.D. (1974) - The anatomy of Saurosuchus galilei and the relationships of the rauisuchid thecodonts. Bulletin of the Museum of Comparative Zoology, 146: 317-362.
Sues H.-D., Schoch R.R., Sobral G. & Irmis R.B. (2020) - A new archosauriform reptile with distinctive teeth from the Middle Triassic (Ladinian) of Germany. Journal of Vertebrate Paleontology, e1764968 (14 pp.). Doi: 10.1080/02724634.2020.1764968.
Sulej T. (2005) - A new rauisuchian reptile (Diapsida: Archosauria) from the Late Triassic of Poland. Journal of Vertebrate Paleontology, 25: 78-86.
Tichy G. (1998) - Ein Neufund von Neusticosaurus toeplitschi (Nopcsa, 1928) (Reptilia, Sauropterygia) aus den Partnachschichtender Gailtaler Alpen, Karnten. Carinthia II, 108: 519-530.
Wachtler M. (2018) - The marine reptile Neusticosaurus from the Eastern Alps. In: Perner T. & Wachtler M. (eds.), Some new and exciting Triassic Archosauria from the Dolomites (Northern Italy). Dolomythos Museum - Oregon Institute of Geological Research. Pp. 47-57.
Walker A. D. (1964) - Triassic reptiles from the Elgin area Ornithosuchus and the origin of Carnosaurs. Philosophical Transactions of the Royal Society of London, 744: 53-134.
Warch A. (1979) - Perm und Trias der nördlichen Gailtaler Alpen. Carinthia II, Sonderhefte 35: 1-111.
Wild R. (1973) - Die Triasfauna der Tessiner Kalkalpen. XXIII. Tanystropheus longobardicus (Bassani) (Neue Ergebnisse). Schweizerische Paläontologische Abhandlungen, 95: 1-162.
Zapfe H. & König H. (1980) - Neue Reptilienfunde aus der Mitteltriasder Gailtaler Alpen (Kärnten, Österreich). Sitzungsberichteder Österreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abt. I, 189: 6582.
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- The matrix used to test the affinities of Carinthiasuchus kandutschi among the archosauriforms. The new taxon has been included in the data set of Desojo & Rauhut (2024), which is an updated version of the phylogenetic matrix of Nesbitt (2011).
- Fig. 1 - High resolution photograph of the row of articulated mid-distal caudal vertebrae.
- Fig. 2- The whole reduced strict consensus of the shortest trees found by the phylogenetic analysis after a posteriori pruning of six "wildcard" terminal taxa (see text). Number at nodes indicate the Bremer support values.
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Accepted 2026-04-09
Published 2026-06-08


